Modern avian lineages evolved and radiated during the Cenozoic Era (66 Ma–present) following the end-Cretaceous extinction. Despite a wealth of isolated eggshells and nest structures, the global oological record remains unevenly studied. This review integrates the scattered literature on Cenozoic oological remains to produce the first comprehensive synthesis of their distribution, structure, palaeobiogeography, and palaeobiological significance. This record reconstructs temporal patterns in eggshell microstructure, nesting ecology, and biogeography. Methodological advances in microscopy, electron backscatter diffraction, isotopic geochemistry, and biomolecular preservation have enabled and refined taxonomic attributions and ecological interpretations of oological material. This synthesis highlights key gaps—particularly in Africa, Antarctica, and Oceania—and outlines research priorities aimed at linking palaeontology, molecular biology, conservation, de-extinction, and Earth-system sciences to better understand the evolutionary trajectory of avian reproduction. Furthermore, this evolving avian fossil record offers a critical foundation for reconstructing past ecosystems and informing efforts to conserve biodiversity into the future
BarrowcloughGFCracraftJKlickaJ, et al.How Many Kinds of Birds Are There and Why Does It Matter?PLOS ONE2016; 11: e0166307.
3.
WellnhoferP.A short history of pterosaur research. Zitteliana B2008; 28: 7–19.
4.
WellnhoferP.A short history of research on Archaeopteryx and its relationship with dinosaurs. In: MoodyRTJBuffetautENaishD, et al. (eds) Dinosaurs and Other Extinct Saurians: A Historical Perspective. London: The Geological Society of London2010, pp. 237–250.
5.
NaishD. The fossil record of bird behaviour. J Zool2014; 292: 268–280.
6.
MainwaringMCMedinaITobalskeBW, et al.The evolution of nest site use and nest architecture in modern birds and their ancestors. Philos Trans R Soc B2023; 378: 20220143.
7.
SchweitzerMHJacksonFDChiappeLM, et al.Late Cretaceous avian eggs with embryos from Argentina. J Vertebr Paleontol2002; 22: 191–195.
8.
TambussiCPNoriegaJI. The Fossil Record of Condors (Ciconiiformes: Vulturidae) in Argentina. Smithson Contrib Paleobiology1999; 89: 177–184.
9.
SánchezMarco A. Tertiary avian localities of Spain. Acta Univ Carol Geol1996; 39: 719–732.
JadwiszczakP. Penguin past: The current state of knowledge. Pol Polar Res2009; 30: 3–28.
12.
AngstDLécuyerCAmiotR, et al.Isotopic and anatomical evidence of an herbivorous diet in the Early Tertiary giant bird Gastornis. Implications for the structure of Paleocene terrestrial ecosystems. Naturwissenschaften2014; 101: 313–322.
13.
AgnolinFL. Unexpected diversity of ratites (Aves, Palaeognathae) in the early Cenozoic of South America: palaeobiogeographical implications. Alcheringa Australas J Palaeontol2016; 41: 101–111.
14.
Vieco-GálvezDCastroIMorelPCH, et al.The eggshell structure in apteryx: form, function, and adaptation. Ecol Evol2021; 11: 3184–3202.
15.
MikhailovKEZelenkovN. The late Cenozoic history of the ostriches (Aves: Struthionidae), as revealed by fossil eggshell and bone remains. Earth-Sci Rev2020; 208: 103270.
16.
SánchezMarco A. Catálogo paleornitológico del Terciario ibérico y balear. Estud Geológicos1999; 55: 163–171.
17.
MontanariS. Cracking the egg: the use of modern and fossil eggs for ecological, environmental and biological interpretation. R Soc Open Sci2018; 5: 180006.
18.
ChoiSHauberMELegendreLJ, et al.Microstructural and crystallographic evolution of palaeognath (Aves) eggshells. eLife2023; 12: e81092.
19.
LazzeriniNLécuyerCAmiotR, et al.Oxygen isotope fractionation between bird eggshell calcite and body water: application to fossil eggs from Lanzarote (Canary Islands). Sci Nat2016; 103: 1–5.
20.
JohnsonBJMillerGHFogelML, et al.The determination of late Quaternary paleoenvironments at Equus Cave, South Africa, using stable isotopes and amino acid racemization in ostrich eggshell. Palaeogeogr Palaeoclimatol Palaeoecol1997; 136: 121–137.
21.
GiardinaMANemeGAGilAF. Rheidae Egg Human Exploitation and Stable Isotopes: Trends from West Central Argentina. Int J Osteoarchaeol2014; 24: 166–186.
22.
MikhailovK. Fossil and recent eggshell in amniotic vertebrates: Fine structure, comparative morphology and classification. London: Palaeontological Association, 1997.
23.
ChoiSParkYMoreno-AzanzaM. Protocol for electron backscatter diffraction (EBSD) analysis of fossil eggshells. J Vertebr Paleontol2023; 43: e2363210.
24.
MingardKP. The use of EBSD for analysis of additively manufactured materials. 152, National Physical Laboratory, 7 February 2023.
25.
WangFEchlinMPTaylorAA, et al.Electron backscattered diffraction using a new monolithic direct detector: High resolution and fast acquisition. Ultramicroscopy2021; 220: 113160.
26.
GouldenJTrimbyPBewickA. The Benefits and Applications of a CMOS-based EBSD Detector. Microsc Microanal2018; 24: 1128–1129.
27.
WiniarskiBGholiniaAMingardK, et al.Correction of artefacts associated with large area EBSD. Ultramicroscopy2021; 226: 113315.
28.
FowlerJERugglesTJCillessenDE, et al.High-Throughput Microstructural Characterization and Process Correlation Using Automated Electron Backscatter Diffraction. Integrating Mater Manuf Innov2024; 13: 641–655.
29.
HirschKFPackardMJ. Review of Fossil Eggs and Their Shell Structure. Scanning Microsc1987; 1: 383–400.
30.
JacksonFDSchweitzerMHSchmittJG. Dinosaur eggshell study using scanning electron microscopy. Scanning J Scanning Microsc2002; 24: 217–223.
31.
NewburyDERitchieNWM. Electron-Excited X-ray Microanalysis by Energy Dispersive Spectrometry at 50: Analytical Accuracy, Precision, Trace Sensitivity, and Quantitative Compositional Mapping. Microsc Microanal2019; 25: 1075–1105.
32.
SharpWDTryonCANiespoloEM, et al.230Th/U burial dating of ostrich eggshell. Quat Sci Rev2019; 219: 263–276.
33.
YangS-YJiangQMChenZ-Y, et al.Electron Probe Microanalysis In Geosciences: Analytical Procedures and Recent Advances. At Spectrosc2022; 43: 186–200.
34.
GrealyAMillerGHPhillipsMJ, et al.Molecular exploration of fossil eggshell uncovers hidden lineage of giant extinct bird. Nat Commun2023; 14: 914.
35.
GrealyAPhillipsMMillerG, et al.Eggshell palaeogenomics: Palaeognath evolutionary history revealed through ancient nuclear and mitochondrial DNA from Madagascan elephant bird (Aepyornis sp.) eggshell. Mol Phylogenet Evol2017; 109: 151–163.
36.
SchweitzerMHChiappeLGarridoAC, et al.Molecular preservation in Late Cretaceous sauropod dinosaur eggshells. Proc R Soc B Biol Sci2005; 272: 775–784.
37.
LegendreLJChoiSClarkeJA. The diverse terminology of reptile eggshell microstructure and its effect on phylogenetic comparative analyses. J Anat2022; 241: 641–666.
38.
PoropatSFTosoliniA-MPBeestonSL, et al.Fossilized gut contents elucidate the feeding habits of sauropod dinosaurs. Curr Biol2025; 35: 2597-2613.e7.
39.
ChiappeLMCoriaACDingusL, et al.Sauropod dinosaur embryos from the Late Cretaceous of Patagonia. Nature1998; 396: 258–261.
40.
SatoTChengYNWuX-C, et al.A pair of shelled eggs inside a female dinosaur. Science2005; 308: 375–376.
41.
BiSAmiotRPeyre De FabrèguesC, et al.An oviraptorid preserved atop an embryo-bearing egg clutch sheds light on the reproductive biology of non-avialan theropod dinosaurs. Sci Bull2021; 66: 947–954.
42.
OliveiraCEMSantucciRMAndradeMB, et al.Crocodylomorph eggs and eggshells from the Adamantina Formation (Bauru Group), Upper Cretaceous of Brazil. Palaeontology2011; 54: 309–321.
43.
CadenaE-AParra-RugeMLParra-RugeJ de D, et al.A gravid fossil turtle from the Early Cretaceous reveals a different egg development strategy to that of extant marine turtles. Palaeontology2019; 62: 533–545.
44.
KeYWuRZelenitskyDK, et al.A large and unusually thick-shelled turtle egg with embryonic remains from the Upper Cretaceous of China. Proc R Soc B Biol Sci2021; 288: 20211239.
45.
Grellet-TinnerGNorellM. An avian egg from the Campanian of Bayn Dzak, Mongolia. J Vertebr Paleontol2002; 22: 719–721.
46.
ZelenitskyDKModestoSP. New information on the eggshell of ratites (Aves) and its phylogenetic implications. Can J Zool2003; 81: 962–970.
47.
LegendreLJRubilar-RogersDMusserGM, et al.A giant soft-shelled egg from the Late Cretaceous of Antarctica. Nature2020; 583: 411–414.
48.
HarrisonT.Fossil bird eggs from the Pliocene of Laetoli, Tanzania: Their taxonomic and paleoecological relationships. J Afr Earth Sci2005; 41: 289–302.
49.
HarrisonTMsuyaCP. Fossil struthionid eggshells from Laetoli, Tanzania: taxonomic and biostratigraphic significance. J Afr Earth Sci2005; 41: 303–315.
50.
HaywardJLDicksonKMGambleSR, et al.Eggshell taphonomy: environmental effects on fragment orientation. Hist Biol2011; 23: 5–13.
51.
WilsonHMHeckCTVarricchioDJ, et al.Evaluating deformation in Spheroolithus dinosaur eggs from Zhejiang, China. Hist Biol2014; 26: 173–182.
52.
JanssenJDMutchGWHaywardJL. Taphonomic Effects of High Temperature on Avian Eggshell. Palaios2011; 26: 658–664.
53.
McGrathACVarricchioDJHaywardJL. Taphonomic assessment of material generated by an Arboreal Nesting Colony of Great Blue Herons (Ardea herodias). Hist Biol2021; 33: 1781–1795.
54.
PortugalSJMaurerGCasseyP. Eggshell Permeability: A Standard Technique for Determining Interspecific Rates of Water Vapor Conductance. Physiol Biochem Zool2010; 83: 1023–1031.
55.
WittonMPHabibMB. On the Size and Flight Diversity of Giant Pterosaurs, the Use of Birds as Pterosaur Analogues and Comments on Pterosaur Flightlessness. PLOS ONE2010; 5: e13982.
56.
VarricchioDJJacksonFD. Reproduction in Mesozoic birds and evolution of the modern avian reproductive mode. The Auk2016; 133: 654–684.
57.
TanakaKZelenitskyDKLüJ, et al.Incubation behaviours of oviraptorosaur dinosaurs in relation to body size. Biol Lett2018; 14: 20180135.
58.
Mourer-ChauviréC. Paleogene avian localities of France. Acta Univ Carol Geol1996; 39: 567–598.
59.
DonaireMLópez-MartínezN. Porosity of late paleocene ornitholithus eggshells (Tremp Fm, south-central Pyrenees, Spain): palaeoclimatic implications. Palaeogeogr Palaeoclimatol Palaeoecol2009; 279: 147–159.
60.
AngstDAmiotRBuffetautE, et al.Diet and climatic context of giant birds inferred from δ13Cc and δ18Oc values of Late Palaeocene and Early Eocene eggshells from southern France. Palaeogeogr Palaeoclimatol Palaeoecol2015; 435: 210–221.
61.
AubryM-POudaKDupuisC, et al.The Global Standard Stratotype-section and Point (GSSP) for the base of the Eocene Series in the Dababiya section (Egypt). Episodes J Int Geosci2007; 30: 271–286.
62.
Mourer-ChauviréC. Les oiseaux fossiles des phosphorites du quercy (éocène supérieur a oligocène supérieur): Implications paléobiogéographiques. Geobios1982; 15: 413–426.
63.
HoudeP. Paleognathous Birds from the Early Tertiary of the Northern Hemisphere. Publ Nuttall Ornithol Club1988; 22: 1–148.
64.
TylerCSimkissK. Studies on egg shells. XII.—Some changes in the shell during incubation. J Sci Food Agric1959; 10: 611–615.
65.
Grellet-TinnerGDykeGJ. The eggshell of the Eocene bird Lithornis. Acta Palaeontol Pol2005; 50: 831–835.
66.
MeiseW. Verhalten der Straussartigen Vogel und Monophylie der Ratite. Proc XIII Int Ornithol Congr1963; 1: 15–125.
67.
LeggittVL. Preliminary Report on the stratigraphic setting and microstructure of avian eggshell fragments from the Calico Mountains, Barstow Formation, Mojave Desert, California. In: Between the Basins: Exploring the Western Mojave and Southern Basin and Range Province. Fullerton, CA: California State University, Desert Studies Consortium, pp. 31–57.
68.
McGrewPO. An Eocene flamingo nesting area, Sweetwater County, Wyoming. Natl Geogr Res Rep1980; 12: 473–478.
69.
McGrewPOFeducciaA. A Preliminary Report on a Nesting Colony of Eocene Birds. In: 25th Field Conference Wyoming Geological Association Guidebook. 1973, pp. 163–165.
70.
GarciaGMourer-ChauviréCAdaciM, et al.First discovery of avian egg and bone remains (Presbyornithidae) from the Gour Lazib (Eocene, Algeria). J Afr Earth Sci2020; 162: 103666.
71.
EricsonPGP. Systematic relationships of the palaeogene family Presbyornithidae (Aves: Anseriformes). Zool J Linn Soc1997; 121: 429–483.
72.
MayrG. Pelagornithidae, Gastornithidae, and Crown Group Galloanseres. In: MayrG (ed) Paleogene Fossil Birds. Cham: Springer International Publishing, pp. 43–72.
73.
KurochkinENDykeGJ. A large collection of Presbyornis (Aves, Anseriformes, Presbyornithidae) from the late Paleocene and early Eocene of Mongolia. Geol J2010; 45: 375–387.
74.
De PietriVLScofieldRPZelenkovN, et al.The unexpected survival of an ancient lineage of anseriform birds into the Neogene of Australia: the youngest record of Presbyornithidae. R Soc Open Sci2016; 3: 150635.
75.
Grellet-TinnerGChiappeLNorellM, et al.Dinosaur eggs and nesting behaviors: A paleobiological investigation. Palaeogeogr Palaeoclimatol Palaeoecol2006; 232: 294–321.
76.
FernándezMSGarciaRAFiorelliL, et al.A large accumulation of avian eggs from the Late Cretaceous of Patagonia (Argentina) reveals a novel nesting strategy in Mesozoic birds. PLOS ONE2013; 8: e61030.
77.
KohringRHirschKF. Crocodilian and avian eggshells from the Middle Eocene of the Geiseltal, Eastern Germany. J Vertebr Paleontol1996; 16: 67–80.
78.
MayrG. An updated review of the middle Eocene avifauna from the Geiseltal (Germany), with comments on the unusual taphonomy of some bird remains. Geobios2020; 62: 45–59.
79.
GrundlerMCRaboskyDL. Rapid increase in snake dietary diversity and complexity following the end-Cretaceous mass extinction. PLoS Biol2021; 19: e3001414.
80.
HirschKFKihmAJZelenitskyDK. New eggshell of ratite morphotype with predation marks from the Eocene of Colorado. J Vertebr Paleontol1997; 17: 360–369.
81.
JacksonFDVarricchioDJCorsiniJA. Avian eggs from the Eocene Willwood and Chadron formations of Wyoming and Nebraska. J Vertebr Paleontol2013; 33: 1190–1201.
82.
HirschKFBowlesJ. Early Eocene Crane-Like eggs? In: Proceedings of the Crane Workshop. Colorado: Colorado State University Printing Service, pp. 211–216.
PanadèsIBlasXGalobartÀArcherM, et al.Australia’s oldest crocodylian eggshell: insights into the reproductive paleoecology of mekosuchines. J Vertebr Paleontol2025; 45: e2560010.
85.
FieldDJ. Big-time insights from a tiny bird fossil. Proc Natl Acad Sci U S A2017; 114: 7750–7752.
86.
HutchinsonDKCoxallHKLuntDJ, et al.The Eocene–Oligocene transition: a review of marine and terrestrial proxy data, models and model–data comparisons. Clim Past2021; 17: 269–315.
87.
CoxallHKPearsonPN. The Eocene–Oligocene Transition. In: WilliamsMHaywoodAMGregoryFJ, et al. (eds) Deep-Time Perspectives on Climate Change: Marrying the Signal from Computer Models and Biological Proxies. Geological Society ofLondon, pp. 351–387.
88.
FarringtonOC. A fossil egg from South Dakota. Publ Field Columbian Mus1899; 35: 193–200.
ChandlerRMWallWP. The first record of bird eggs from the early Oligocene (Orellan) of North America. In: SantucciVLMcClellandL (eds) Proceedings of the 6th Fossil Resource Conference. 2001, pp. 49–54.
91.
SinclairWJ. The Faunas of the Concretionary Zones of the Oreodon Beds, White River Oligocene. Proc Am Philos Soc1924; 63: 94–133.
92.
LawverDRBoydCA. An avian eggshell from the Brule Formation (Oligocene) of North Dakota. J Vertebr Paleontol2018; 38: 1–9.
93.
HasiotisSTBownTMAbstonC. Photoglossary of marine and continental ichnofossils. USGS Numbered Series23, 1994.
94.
WesterholdTMarwanNDruryAJ, et al.An astronomically dated record of Earth’s climate and its predictability over the last 66 million years. Science2020; 369: 1383–1387.
95.
MethnerKCampaniMFiebigJ, et al.Middle Miocene long-term continental temperature change in and out of pace with marine climate records. Sci Rep2020; 10: 7989.
96.
PolissarPJUnoKTPhelpsSR, et al.Hydrologic Changes Drove the Late Miocene Expansion of C4 Grasslands on the Northern Indian Subcontinent. Paleoceanogr Paleoclimatology2021; 36: e2020PA004108.
97.
BrandtA. Über ein grosses fossiles Vogelei aus der Umgebung von Cherson. In: Bulletin de l’Académie Impériale des Sciences de St Pétersbourg. 1873, pp. 158–161.
98.
BrandtA. II.—On a large Fossil Egg from the neighbourhood of Cherson. Ibis1874; 3: 4–7.
99.
PatnaikRSahniACameronD, et al.Ostrich-like eggshells from a 10.1 million-yr-old Miocene ape locality, Haritalyangar, Himachal Pradesh, India. Curr Sci2009; 96: 1485–1495.
100.
TøienØPaganelliCVRahnH, et al.Diffusive resistance of avian eggshell pores. Respir Physiol1988; 74: 345–354.
101.
AttardMRGPortugalSJ. Global diversity and adaptations of avian eggshell thickness indices. Ibis2024; 166: 534–550.
102.
TanakaKZelenitskyDKTherrienF. Eggshell porosity provides insight on evolution of nesting in dinosaurs. PLOS ONE2015; 10: e0142829.
103.
BoardRGScottVD. Porosity of the Avian Eggshell. Am Zool1980; 20: 339–349.
104.
ZelenkovNVKurochkinEN. Birds of Central Asia in the Neogene. In: Ornitologiya v Tsentral’noi Evrazii: Materialy XIII Mezhdunar. Ornitologicheskaya Konferentsiya Severnoi Evrazii. Tezisy Dokladov. Orenburg: Gosudarstvenny Pedagogichesky Universitet, 2010, pp. 133–134.
105.
SenutBPickfordMSégalenL. Neogene desertification of Africa. Comptes Rendus Geosci2009; 341: 591–602.
106.
PickfordMSenutBDauphinY. Biostratigraphy of the Tsondab Sandstone (Namibia) based on gigantic avian eggshells. Géobios1995; 28: 85–98.
107.
SmithNA. Avian fossils from the early Miocene Moghra Formation of Egypt. Ostrich2013; 84: 181–189.
108.
BuffetautE. Dragon’s eggs’ from the ‘Yellow Earth’: The discovery of the fossil ostriches of China. Hist Nat2021; 11: 47–63.
109.
MlíkovskýJ. Early Miocene birds of Djebel Zelten, Libya. Časopis Národního Muzea Řada Přír2003; 172: 114–120.
110.
TessierFHébrardLLappartientJR. Découverte de fragments d’œufs de Psammorniset de Struthio dans le Quaternaire de la presqu’île du Cap Blanc (République Islamique de Mauritanie). Comptes Rendus Académie Sci Paris Sér D; 273: 2418–2421.
111.
TyrbergT.The Late Pleistocene Continental Avian extinction - an evaluation of the fossil evidence. Oryctos2008; 7: 249–269.
112.
LowePR. On some Struthious Remains: 1. Description of some pelvic remains of a large fossil Ostrich Struthio oldawayi sp. nov. from the Lower Pleistocene of Oldaway (Tanganyica Territory). 2. Egg-shell fragments referable to Psammornis and other Struthiones. Ibis1933; 13: 652–658.
113.
BravoAMSevillaPBarroso-BarcenillaF. Avian and crocodilian eggshells from the upper Barremian site of Vadillos-1 (Lower Cretaceous, Cuenca province, Spain). Cretac Res2018; 85: 28–41.
114.
WorthyTHTennysonAJDJonesC, et al.Miocene waterfowl and other birds from central Otago, New Zealand. J Syst Palaeontol2007; 5: 1–39.
115.
TennysonTWorthyTJonesC, et al.Moa’s Ark: Miocene Fossils Reveal the Great Antiquity of Moa (Aves: Dinornithiformes) in Zealandia. Rec Aust Mus2010; 62: 105–114.
116.
ColeTLWoodJR. The ancient DNA revolution: The latest era in unearthing New Zealand’s faunal history. N Z J Zool2018; 45: 91–120.
117.
WilliamsDLGVickers-RichP. Giant fossil eggs fragment from the Tertiary of Australia. Contrib Sci Nat Hist Mus Los Angel Cty1992; 36: 375–378.
118.
Grellet-TinnerG. A re-evaluation of the Snake Dam eggshell Phylogenetic and temporal implications. J Vertebr Paleontol2001; 21: 57.
119.
WorthyTHDegrangeFJHandleyWD, et al.The evolution of giant flightless birds and novel phylogenetic relationships for extinct fowl (Aves, Galloanseres). R Soc Open Sci2017; 4: 170975.
120.
McInerneyPLBloklandJCWorthyTH. Skull morphology of the enigmatic Genyornis newtoni Stirling and Zeitz, 1896 (Aves, Dromornithidae), with implications for functional morphology, ecology, and evolution in the context of Galloanserae. Hist Biol2024; 36: 1093–1165.
121.
HandleyWDWorthyTH. Endocranial anatomy of the giant extinct Australian Mihirung birds (Aves, Dromornithidae). Diversity2021; 13: 124.
122.
SellésALujánÀHCasanovas-VilarI, et al.Exceptionally preserved eggs show philopatric resilience of phasianids to environmental fragmentation during the Middle–Late Miocene transition in the northeastern Iberian Peninsula. Pap Palaeontol2025; 11: e70060.
123.
MarmiJCasanovas-VilarIRoblesJM, et al.The paleoenvironment of Hispanopithecus laietanus as revealed by paleobotanical evidence from the Late Miocene of Can Llobateres 1 (Catalonia, Spain). J Hum Evol2012; 62: 412–423.
124.
ArranzSGCasanovas-VilarIŽliobaitėI, et al.Paleoenvironmental inferences on the Late Miocene hominoid-bearing site of Can Llobateres (NE Iberian Peninsula): An ecometric approach based on functional dental traits. J Hum Evol2023; 185: 103441.
125.
Agustí i BallesterJMoyà SolàSCabrera PérezL. Sinopsis estratigráfica del Neógeno de la fosa del Vallés-Penedés. Paleontol Evolució1983; 57–82.
126.
KohringR. Schildkriten-und Vogeleischalen aus dem Untermiozdin (MN4b) von Dolnice (Tschechien). Doc Naturae1998; 122: 49–57.
127.
Amo-SanjuanOLaplanaC. Primer registro de fragmentos de cáscara de huevo de ave en el Terciario de la Cuenca del Ebro. In: MeléndezGHerreraZDelveneG, et al. (eds) Publicaciones del Seminario de Paleontología de Zaragoza. 2001, pp. 199–204.
128.
HirschKBrayES. Spheroidal eggs -avian and chelonian-from the Miocene and Oligocene of the Western Interior. Hunteria1988; 1: 1–8.
129.
GrossMKohringRSümeczF. Sie singen seit 11,5 Mio. Jahren: Das älteste Singvogelei der Welt. Foss – Z Für Hobbypaläontologen2008; 5: 293–299.
130.
BarkerFKCiboisASchiklerP, et al.Phylogeny and diversification of the largest avian radiation. Proc Natl Acad Sci U S A2004; 101: 11040–11045.
131.
OliverosCHFieldDJKsepkaDT, et al.Earth history and the passerine superradiation. Proc Natl Acad Sci U S A2019; 116: 7916–7925.
132.
MoyleRGOliverosCHAndersenMJ, et al.Tectonic collision and uplift of Wallacea triggered the global songbird radiation. Nat Commun2016; 7: 12709.
133.
MikulášRZasadilB. A fossil bird nest, the find from the Tertiary, Northern Czech Republic. Vesmír2002; 79, 422: 425–426.
134.
MikulášRZasadilB. A probable fossil bird nest, ?Eocavum isp., from the Miocene wood of the Czech Republic. In: RM (ed) Abstract Book. 4th International Bioerosion Workshop. 2004, pp. 49–51.
135.
PoinarGJVoisinC. Bird eggshell in Dominican Amber. Palaeontology2007; 50: 1381–1383.
136.
Iturralde-VinentMAMacPheeRDE. Age and Paleogeographical Origin of Dominican Amber. Science1996; 273: 1850–1852.
137.
Kosmowska-CeranowiczBKulickiCKupryjanowiczJ, et al.Bird traces in Dominican Amber. In: Kosmowska-CeranowiczBGierlowskiWSontagE (eds) The Eocene Climatic Optimum and the formation of the Baltic amber deposits. Gdańsk, Poland: Gdańsk International Fair Co., 2013, pp. 62–63.
138.
Grellet-TinnerGMurelagaXLarrasoañaJC, et al.The First Occurrence in the Fossil Record of an Aquatic Avian Twig-Nest with Phoenicopteriformes Eggs: Evolutionary Implications. PLOS ONE2012; 7: e46972.
139.
MurelagaXAlmarYBeamudB, et al.Nueva localidad fosilífera en el Mioceno Inferior de Bargota (Navarra)(Cuenca del Ebro, Península Ibérica). Geogaceta2006; 40: 171–174.
140.
MartínezGarcía BGonzález PardosMPascualA, et al.Primeros datos paleontológicos de la sección de Loma Negra (Bardenas Reales de Navarra, Mioceno inferior-medio). Geogaceta2013; 54: 63–66.
141.
Suárez HernandoOZuluaga IbargallartuMCMartínezGarcía B, et al.Análisis mineralógico de las arcillas del tránsito Mioceno inferior-medio en la sección Loma Negra (Bardenas Reales de Navarra, Cuenca del Ebro). Geogaceta2016; 60: 111–114.
142.
AmoOAzanzaBSánchezA, et al.First record of ornitoid prismatic eggshells from the Miocene of Spain. In: IV European workshop on vertebrate palaeontology. Albarracín, Teruel, Spain, 1999, pp. 16–17.
143.
AzanzaBAlonso-ZarzaAÁlvarez SierraMA, et al.Los yacimientos de vertebrados continentales del Aragoniense superior (Mioceno medio) de Toril, Cuenca de Calatayud-Daroca. Geo-Temas2004; 6: 271–274.
144.
HuttererRGarcía-TalaveraFSánchez-PintoL. Huevos y huesos de pájaros del Mioceno. en las Islas Salvajes. Rev Acad Canar Cienc2001; 13: 23–29.
145.
MeijerHJM. In het kielzog van Darwin–Ei zoeken op Selvagens. Cranium2009; 26: 48–51.
146.
MassironiMSauroFMeyzenC, et al.Geology of Selvagens. In: Sevilla. Casa de la Ciencia-CSIC de Sevilla: CSIC - Instituto de Recursos Naturales y Agrobiología de Sevilla (IRNAS), 2023.
147.
SnidermanJMKWoodheadJDHellstromJ, et al.Pliocene reversal of late Neogene aridification. Proc Natl Acad Sci U S A2016; 113: 1999–2004.
148.
HallekFOuajaMHallekT, et al.Upper Neogene climate change in northern Africa based on chemical weathering indices and clay mineralogy: a case study of southeastern Tunisia (Gulf of Gabès). J Sediment Environ2024; 9: 317–336.
149.
BuffetautE. The Enigmatic Avian Oogenus Psammornis: A Review of Stratigraphic Evidence - ProQuest. Diversity2022; 14: 123.
150.
JainSRaiNKumarG, et al.Ancient DNA Reveals Late Pleistocene Existence of Ostriches in Indian Sub-Continent. PLOS ONE2017; 12: e0164823.
151.
Hernández-PachecoE. Nuevos yacimientos de vertebrados miocenos y deducciones de orden paleofisiográfico. Asoc Esp Para El Prog Las Cienc1921; 6: 159–170.
152.
Hernández-PachecoE. Los vertebrados terrestres del Mioceno de la Península Ibérica. Mem Real Soc Esp Hist Nat1914; 4: 443–488.
153.
BuchholzHV. Die Höhle eines Spectvogels aus dem Eozän von Arizona, USA (Aves Piciformes). Verhandlungen Naturwissenschaftlichen Ver Hambg1986; 28: 5–25.
154.
García-Talavera CasañasF. Aves gigantes en el Mioceno de Famara (Lanzarote). Rev Acad Canar Cienc Folia Canar Acad Sci1990; 2: 71–79.
155.
MayrG. Two-phase extinction of “Southern Hemispheric” birds in the Cenozoic of Europe and the origin of the Neotropic avifauna. Palaeobiodiversity Palaeoenvironments2011; 91: 325–333.
156.
CasañasFGT. Aves gigantes en el Mioceno de Famara (Lanzarote). Rev Acad Canar Cienc Folia Canar Acad Sci1990; 2: 71–79.
157.
BibiFHillABeechM, et al.Late Miocene Fossils from the Baynunah Formation, United Arab Emirates. In: ForteliusMWangXFlynnL (eds). Columbia University Press, pp. 583–594.
158.
CohenASDuARowanJ, et al.Plio-Pleistocene environmental variability in Africa and its implications for mammalian evolution. Proc Natl Acad Sci U S A2022; 119: e2107393119.
159.
LowePRBateDMA. Struthious remains from China and Mongolia with description of Struthio wimani, Struthio anderssoni and Struthio mongolicus spp. nov. Palaeontol Sin1931; 6: 1–40.
160.
BlinkhornJAchyuthanHPetragliaMD. Ostrich expansion into India during the Late Pleistocene: Implications for continental dispersal corridors. Palaeogeogr Palaeoclimatol Palaeoecol2015; 417: 80–90.
161.
StidhamTA. Extinct ostrich eggshell (Aves: Struthionidae) from the Pliocene Chiwondo Beds, Malawi: implications for the potential biostratigraphic correlation of African Neogene deposits. J Hum Evol2004; 46: 489–496.
162.
KurochkinENKuzminYVAntoshchenko-OlenevIV, et al.The timing of ostrich existence in Central Asia: AMS 14C age of eggshells from Mongolia and southern Siberia (a pilot study). Nucl Instrum Methods Phys Res Sect B Beam Interact Mater At2010; 268: 1091–1093.
MillerGHFogelMLMageeJW, et al.Ecosystem Collapse in Pleistocene Australia and a Human Role in Megafaunal Extinction. Science2005; 309: 287–290.
165.
MillerGHMageeJWJohnsonBJ, et al.Pleistocene extinction of genyornis newtoni: human impact on australian megafauna. Science1999; 283: 205–208.
166.
MillerGHFogelMLMageeJW, et al.Disentangling the impacts of climate and human colonization on the flora and fauna of the Australian arid zone over the past 100 ka using stable isotopes in avian eggshell. Quat Sci Rev2016; 151: 27–57.
167.
WilliamsDLG. Genyornis eggshell (Dromornithidae; Aves) from the Late Pleistocene of South Australia. Alcheringa1981; 5: 133–140.
168.
BirdMITurneyCSMFifieldLK, et al.Radiocarbon dating of organic- and carbonate-carbon in Genyornis and Dromaius eggshell using stepped combustion and stepped acidification. Quat Sci Rev2003; 22: 1805–1812.
169.
Grellet-TinnerGSpoonerNAWorthyTH. Is the “Genyornis” egg of a mihirung or another extinct bird from the Australian dreamtime?Quat Sci Rev2016; 133: 147–164.
170.
ChanNR. Does size variation in Genyornis newtoni (Aves, Dromornithidae) encompass eggshell safety limits?J Vertebr Paleontol2014; 34: 976–979.
171.
MillerGHFogelMLMageeJW, et al.The Genyornis egg: A commentary on Grellet-Tinner et al., 2016. Quat Sci Rev2017; 161: 123–127.
172.
DemarchiBStillerJGrealyA, et al.Ancient proteins resolve controversy over the identity of Genyornis eggshell. Proc Natl Acad Sci U S A2022; 119: e2109326119.
173.
SamondsKECrowleyBERasolofomananaTRN, et al.A new late Pleistocene subfossil site (Tsaramody, Sambaina basin, central Madagascar) with implications for the chronology of habitat and megafaunal community change on Madagascar’s Central Highlands. J Quat Sci2019; 34: 379–392.
174.
BalanoffAMRoweT. Osteological description of an embryonic skeleton of the extinct elephant bird, Aepyornis (Palaeognathae: Ratitae). J Vertebr Paleontol; 27: 1–53.
175.
GoodmanSMRaherilalaoMJMuldoonK. Bird fossils from Ankilitelo Cave: inference about Holocene environmental changes in Southwestern Madagascar. Zootaxa2013; 3750: 534–548.
176.
BurkhardS. Fossile Vogeleischalen aus dem Pleistozän von Weimar-Ehringsdorf. Abh Zent Geol Inst1975; 23: 153–159.
177.
KahlkeH-D. Das Pleistozän von Taubach bei Weimar. Quartärpaläontologie1977; 2: 1–509.
178.
RosasAHuguetRPérez-GonzálezA, et al.The “Sima del Elefante” cave site at Atapuerca (Spain). Estud Geológicos2006; 62: 327–348.
179.
MarquetaMHuguetRNúñez-LahuertaC. Accumulation agents and bird assemblages: The case of the TE9d level at Sima del Elefante (Sierra de Atapuerca, Spain). Int J Osteoarchaeol2023; 33: 652–667.
180.
Núñez-LahuertaCMoreno-AzanzaMPérez-PueyoM, et al.Shearwater Eggs in Lobos 3, a Holocene Site of Fuerteventura (Canary Islands). Diversity2023; 15: 144.
181.
Núñez-LahuertaCMoreno AzanzaMCuenca-BescósG. Avian eggshell remains in the human bearing level TD6 of the Gran Dolina site (Early Pleistocene, Atapuerca, Spain). Hist Biol2021; 33: 660–671.
182.
Núñez-LahuertaCGalánJCuenca-BescósG, et al.A bird assemblage across the MIS 9/8 boundary: The Middle Pleistocene of Galería (Atapuerca). Quat Sci Rev2022; 293: 107708.
183.
OlsonSL. Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C, 4-7 June 1996. Smithson Contrib Paleobiology1999; 1–344.
184.
OlsonSLWingateDBHeartyPJ, et al.Prodromus of Vertebrate Paleontology and Geochronology of Bermuda. In: AlcoverJABoverP (eds) Proceedings of the International Symposium “Insular Vertebrate Evolution: the Palaeontological Approach”. Monografies de la Societat d’Història Natural de les Balears. 2005, pp. 113–128.
185.
OlsonSLHeartyPJ. Probable extirpation of a breeding colony of Short-tailed Albatross (Phoebastria albatrus) on Bermuda by Pleistocene sea-level rise. Proc Natl Acad Sci U S A2003; 100: 12825–12829.
186.
OlsonSLHeartyPJ. Fossilized egg indicates probable breeding of Brown Pelican (Pelecanus occidentalis) on Bermuda in the Middle Pleistocene. Proc Biol Soc Wash2013; 126: 169–177.
187.
WoodCA. The fossil eggs of Bermudan birds. Ibis1923; 65: 193–207.
188.
EmslieSD. Age and diet of fossil California Condors in Grand Canyon, Arizona. Science1987; 237: 768–770.
189.
GaoYSalvatoreMCXuQ, et al.The occupation history of the longest-dwelling Adélie penguin colony reflects Holocene climatic and environmental changes in the Ross Sea, Antarctica. Quat Sci Rev2022; 284: 107494.
190.
YangLGaoYXuQ, et al.Specific occupation of penguins under Neoglacial cooling on the Scott Coast, Antarctica. Quat Sci Rev2021; 264: 107010.
191.
MillarCDSubramanianSHeupinkTH, et al.Adélie penguins and temperature changes in Antarctica: a long-term view. Integr Zool2012; 7: 113–120.
192.
GuthrieDAThomasHWKennedyGL. A New Species of Extinct Late Pleistocene Puffin (Aves: Alcidae) from the Southern California Channel Islands. In: Proceedings of the Fifth California Islands Symposium. Santa Barbara: Santa Barbara Museum of Natural History, 2002, pp. 525–530.
193.
CruzJAMoreno-FloresOCoronaM E, et al.The first American occurrence of Phoenicopteridae fossil egg and its palaeobiogeographical and palaeoenvironmental implications. Hist Biol2024; 36: 2051–2058.
194.
del CampoRM. Huevos subfósiles de grulla en el Valle de México (1). An Inst Biol1944; 15: 312–318.
195.
ScottJJRenautRWOwenRB. Impacts of flamingos on saline lake margin and shallow lacustrine sediments in the Kenya Rift Valley. Sediment Geol2012; 277: 32–51.
196.
KsepkaDTBertelliSGianniniNP. The phylogeny of the living and fossil Sphenisciformes (penguins). Cladistics2006; 22: 412–441.
197.
KsepkaDTAndoT. Penguins past, present, and future: Trends in the evolution of the Sphenisciformes. In: DykeGKaiserG (eds) Living Dinosaurs: The Evolutionary History of Modern Birds. John Wiley & Sons, Ltd, 2011, pp. 155–186.
198.
WorthyTHGrant‐MackieJA. Late‐Pleistocene avifaunas from Cape Wanbrow, Otago, South Island, New Zealand. J R Soc N Z2003; 33: 427–485.
199.
KuhnBFCarlsonKJHopleyPJ, et al.Identification of fossilized eggshells from the Taung hominin locality, Taung, Northwest Province, South Africa. Palaeontol Electron2015; 18: 1–16.
200.
BergerLRClarkeRJ. Eagle involvement in accumulation of the Taung child fauna. J Hum Evol1995; 29: 275–299.
201.
Lee-ThorpJASponheimerMLuytJ. Tracking changing environments using stable carbon isotopes in fossil tooth enamel: an example from the South African hominin sites. J Hum Evol2007; 53: 595–601.
202.
McGrawWSCookeCShultzS. Primate remains from African crowned eagle (Stephanoaetus coronatus,) nests in Ivory Coast’s Tai Forest: Implications for primate predation and early hominid taphonomy in South Africa. Am J Phys Anthropol2006; 131: 151–165.
203.
TyrbergT. Pleistocene birds of the Palearctic: a catalogue. Cambridge, Massachusetts: Nuttall Ornithological Club, 1998.
204.
MedinaMELópezMLCamposMR, et al.Pit-houses, seasonality, and subsistence resources: an essay from Boyo Paso 2 (ca. 900–700 BP, Sierras of Córdoba, Argentina). Archaeol Anthropol Sci2020; 12: 119.
205.
ApolinaireETurnesL. Diferenciación específica de Rheidos a partir de fragmentos de cáscaras de huevo. Su aplicación en sitios arqueológicos del Holoceno Tardío. In: BerónMLunaLBonomoM, et al. (eds) Mamül Mapu: pasado y presente desde la Arqueología Pampeana. Ayacucho, Ediciones del Espinillo: Ediciones del Espinillo, 2010, pp. 253–260.
206.
MedinaMEPicassoMBJCamposMR, et al.Tarsometatarsus, eggshells, and the species level identification of large‐sized flightless birds from Boyo Paso 2 (Sierras of Córdoba, Argentina). Int J Osteoarchaeol2019; 29: 584–594.
207.
AmeghinoF. Obras completas y correspondencia científica de Florentino Ameghino. La Plata, Argentina: Taller de impresiones oficiales, 1917. Epub ahead of print 1917. DOI: 10.5962/bhl.title.60823.
208.
GillBJ. Thickness histograms of Holocene fossil eggshell fragments indicate diversity and relative abundance of moas (Aves: Dinornithiformes) at North Island sites. N Z J Zool2022; 49: 143–165.
209.
GillBJ. A Catalogue of Moa Eggs (aves: Dinornithiformes). Rec Auckl Mus2006; 43: 55–80.
210.
HuynenLGillBJMillarCD, et al.Ancient DNA reveals extreme egg morphology and nesting behaviour in New Zealand’s extinct moa. Proc Natl Acad Sci U S A2010; 107: 16201–16206.
211.
WorthyTH. Fossil deposits in the Hodge Creek Cave System, on the northern foothills of Mt Arthur, Nelson, South Island, New Zealand. Notornis1997; 44: 111–124.
212.
WoodRJ. Moa (Aves: Dinornithiformes) nesting material from rockshelters in the semi-arid interior of South Island, New Zealand. J R Soc N Z2008; 38: 115–129.
213.
WorthyTHHoldawayRN. Quaternary fossil faunas from caves on Mt Cookson, North Canterbury, South Island, New Zealand. J R Soc N Z1995; 25: 333–370.
214.
WorthyT. Quaternary fossil faunas of Otago, South Island, New Zealand. J R Soc N Z1998; 28: 421–521.
215.
BoastA. Palaeoecology and Ancient DNA of the Kākāpō (Strigops habroptilus) [dissertation]. Doctoral Dissertation, University of Auckland, 2020.
216.
WoodJR. Subfossil Kākāpō (Strigops habroptilus) remains from near Gibraltar Rock, Cromwell Gorge, Central Otago, New Zealand. Notornis2006; 53: 191.
217.
EmslieSD. Ancient Adélie penguin colony revealed by snowmelt at Cape Irizar, Ross Sea, Antarctica. Geology2021; 49: 145–149.
218.
OskamCLHaileJMcLayE, et al.Fossil avian eggshell preserves ancient DNA. Proc R Soc B Biol Sci2010; 277: 1991–2000.
219.
RowleyGD. On the egg of Aepyornis, the colossal bird of Madagascar. Proc Zool Soc Lond1867; 1867: 892–895.
220.
GrandidierA. V.—Observations on the Egg-beds of Æpyornis. Ibis1868; 10: 65–68.
221.
MonnierL. Paléontologie de Madagascar. VII. Les Aepyornis. Ann Paléontol1913; 8: 125.
222.
RoweTCarlsonWBottorffW. Thrinaxodon-Digital Atlas of the Skull. In: University of Texas Press. 1993, pp. 871–874.
223.
KetchamRACarlsonWD. Acquisition, optimization and interpretation of X-ray computed tomographic imagery: applications to the geosciences. Comput Geosci2001; 27: 381–400.
224.
AllentoftMERawlenceNJ. Moa’s Ark or volant ghosts of Gondwana? Insights from nineteen years of ancient DNA research on the extinct moa (Aves: Dinornithiformes) of New Zealand. Ann Anat - Anat Anz2012; 194: 36–51.
225.
JarvisEDMirarabSAbererAJ, et al.Whole-genome analyses resolve early branches in the tree of life of modern birds. Science2014; 346: 1320–1331.
226.
ClaramuntSCracraftJ. A new time tree reveals Earth history’s imprint on the evolution of modern birds. Sci Adv2015; 1: e1501005.
227.
ViannaJAFernandesFANFrugoneMJ, et al.Genome-wide analyses reveal drivers of penguin diversification. Proc Natl Acad Sci U S A2020; 117: 22303–22310.
228.
StillerJFengSChowdhuryA-A, et al.Complexity of avian evolution revealed by family-level genomes. Nature2024; 629: 851–860.
229.
CloutierASacktonTBGraysonP, et al.Whole-Genome Analyses Resolve the Phylogeny of Flightless Birds (Palaeognathae) in the Presence of an Empirical Anomaly Zone. Syst Biol2019; 68: 937–955.
230.
SacktonTBGraysonPCloutierA, et al.Convergent regulatory evolution and loss of flight in paleognathous birds. Science2019; 364: 74–78.
231.
HarshmanJBraunELBraunMJ, et al.Phylogenomic evidence for multiple losses of flight in ratite birds. Proc Natl Acad Sci U S A2008; 105: 13462–13467.
232.
PressleeSWilsonJWoolleyJ, et al.The identification of archaeological eggshell using peptide markers. STAR Sci Technol Archaeol Res2017; 3: 89–99.
233.
ProndvaiEBotfalvaiGSteinK, et al.Collection of the thinnest: A unique eggshell assemblage from the Late Cretaceous vertebrate locality of Iharkút (Hungary). Cent Eur Geol2017; 60: 73–133.
234.
StoddardMCYongEHAkkaynakD, et al.Avian egg shape: Form, function, and evolution. Science2017; 356: 1249–1254.
235.
Englert DuursmaDGallagherRVPriceJJ, et al.Variation in avian egg shape and nest structure is explained by climatic conditions. Sci Rep2018; 8: 4141.
ChengCMaZ. Conservation interventions are required to improve bird breeding performance in artificial wetlands. Biol Conserv2023; 278: 109872.
238.
DavidsonNC. How much wetland has the world lost? Long-term and recent trends in global wetland area. Mar Freshw Res2014; 65: 934–941.
239.
DonaldPFPisanoGRaymentMD, et al.The Common Agricultural Policy, EU enlargement and the conservation of Europe’s farmland birds. Agric Ecosyst Environ2002; 89: 167–182.
240.
BarwischIMewesWSchmitz OrnésA, et al.Heavy metal residues in eggshells of Common Cranes (Grus grus) nesting in an agricultural region in north-eastern Germany. J Ornithol2024; 165: 507–520.
241.
MoraMA. Heavy metals and metalloids in egg contents and eggshells of passerine birds from Arizona. Environ Pollut2003; 125: 393–400.
242.
ChenDLetcherRJGauthierLT, et al.Newly discovered methoxylated polybrominated diphenoxybenzenes have been contaminants in the Great Lakes herring gull eggs for thirty years. Environ Sci Technol2012; 46: 9456–9463.
243.
ShapiroB. Pathways to de-extinction: how close can we get to resurrection of an extinct species?Funct Ecol2017; 31: 996–1002.
244.
AllentoftMECollinsMHarkerD, et al.The half-life of DNA in bone: measuring decay kinetics in 158 dated fossils. Proc R Soc B Biol Sci2012; 279: 4724–4733.
245.
SoaresAERNovakBJHaileJ, et al.Complete mitochondrial genomes of living and extinct pigeons revise the timing of the columbiform radiation. BMC Evol Biol2016; 16: 230.
246.
ShapiroBSibthorpeDRambautA, et al.Flight of the Dodo. Science2002; 295: 1683.
247.
IUCNSSC. IUCN SSC Guiding principles on Creating Proxies of Extinct Species for Conservation Benefit. Version 1.0. Gland, Switzerland: IUCN Species Survival Commission, 2016.
248.
JørgensenD. Reintroduction and De-extinction. BioScience2013; 63: 719–720.
249.
GenovesiPSimberloffD. “De-extinction” in conservation: Assessing risks of releasing “resurrected” species. J Nat Conserv2020; 56: 125838.
250.
BuffetautEAngstD. Stratigraphic distribution of large flightless birds in the Palaeogene of Europe and its palaeobiological and palaeogeographical implications. Earth-Sci Rev2014; 138: 394–408.
251.
MikhailovKE. Avian egg-shells: an atlas of scanning electron micrographs. British Ornitologist’s Club Occasional Publications, 1997.
252.
MlíkovskýJ. Cenozoic Birds of the World, Part 1: Europe. Part 1: Europe. Ninox Press, 2002.
253.
MörsT. Wirbeltiere aus dem Miozän von Homberg/Ohm (Vogelsberg, Hessen). Geol Jahrb Hess2010; 136: 109–119.
254.
LeggittVLBuchheimPHBiaggiRE. A new Presbyornis (Aves: Anseriformes) nesting site with abundant eggshell: Eocene fossil lake, Lincoln County, Wyoming. J Vertebr Paleontol1998; 18: 58A.
255.
LeggittVLBuchheimPHBiaggiRE. The Stratigraphic Setting of Three Presbyornis Nesting Sites: Eocene Fossil Lake, Lincoln County, Wyoming. National Park Service PaleontologicalResearch1998; 3: 61–68.
256.
BravoAMYébenesAAbadHM. Cáscaras de huevo de tipo Aepyornis del Mioceno Superior de Alicante (España). Aproximación Filogenética Rev Esp Paleontol2009; 24: 47–58.
257.
Mourer-ChauviréC. Pliocene avian localities of France. Acta Univ Carol Geol1996; 39: 613–618.
258.
SenutB. Fossil ratite eggshells: a useful tool for Cainozoic biostratigraphy in Namibia. Commun Geol Surv Namib2000; 12: 367–373.
SahniAkumarGBajpaIS, et al.A review of Late Pleistocene ostriches (Struthio sp.) in India. Man Environ1990; 15: 41–7.
261.
SauerEGF. Aepyornithoide Eierschalen aus dem Miozaen und Pliozaen von Anatolien, Tuerkei. Aepyornithoid eggshell fragments from the Miocene and Pliocene of Anatolia, Turkey. Palaeontogr Abt Palaeozoologie-Stratigr1976; 153: 62–115.
262.
MeinPDauphinY. Des coquilles d’oeuf de type Aepyomis dans le bassin de Teruel (Pliocene basal). Esp N J Geol Palaont1995; M: 182–191.
263.
MlíkovskýJ. Tertiary avian localities of Moldavia. In: MlíkovskýJ (ed) Tertiary Avian Localities of Europe. 1996, pp. 685–691.
264.
MlíkovskỳJHesseA. Tertiary avian localities of Germany. Acta Univ Carol Geol1996; 39: 619–647.
265.
Gamonal González-IglesiasA. Nuevos datos tafonómicos sobre el yacimiento de vertebrados continentales de Toril 3A. In: Current Trends in Paleontology and Evolution. Cercedilla, Madrid, Spain: Universidad Complutense de Madrid (UCM)-Instituto de Geociencias (IGEO, CSIC-UCM)-Museo Nacional de Ciencias Naturales (MNCN-CSIC), 2015, pp. 140–143.
266.
HernándezJMGarcía FernándezSCaballeroF, et al.Estudio sedimentológico de la sucesión con nuevos niveles de vertebrados fósiles del Mioceno inferior de Lardero (La Rioja).
267.
MurelagaX. Estudio de las faunas de vertebrados del Mioceno inferior de las Bardenas Reales y áreas colindantes (Cuenca del Ebro, Navarra). Unpublished PhD Thesis, Universidad del País Vasco UPV/EHU, 2000.
268.
Grellet-TinnerGMurelagaXLarrasoanaJC, et al.The first occurrence in the fossil record of an aquatic avian twig-nest with Phoenicopteriformes eggs: evolutionary implications.
269.
BukhsianidzeMKoiavaK. Synopsis of the terrestrial vertebrate faunas from the Middle Kura Basin (Eastern Georgia and Western Azerbaijan, South Caucasus). Acta Palaeontol Pol; 63: 441–46.
270.
MikhailovKEKurochkinEN. The eggshells of Struthioniformes from the Palearctic and its position in the system of views on Ratite evolution. In: The Joint Soviet-Mongolian Palaeontological Expedition (ed) Fossil Reptiles and Birds of Mongolia, Transactions. Nauka, Moscow, 1988, pp. 43–64.
271.
BibiFShabelAKraatzB, et al.New fossil ratite (Aves: Palaeognathae) eggshell discoveries from the Late Miocene Baynunah Formation of the United Arab Emirates, Arabian Peninsula. Palaeontol Electron2006; 9: 1–13.
272.
StewartJR, Beech. The Miocene birds of Abu Dhabi (United Arab Emirates) with a discussion of the age of modern species and genera. Hist Biol2006; 18: 103–113.
273.
LouchartABibiFStewartJR. Birds from the Baynunah Formation. In: Sands of Time: Ancient life in the late Miocene of Abu Dhabi, United Arab Emirates. Cham: Springer International Publishing, 2022, pp. 125–139.
274.
SenutBPickfordM. Fossil eggs and Cenozoic continental biostratigraphy of Namibia. Palaeontol Afr1995; 32: 33–37.
275.
Burchak-AbramovichNIKonkovaNI. Nakhodki iskopaemykh strausov v MSSR i drugikh punktakh SSSR. In: Paleontologiya, geologiya i poleznye iskopaemye Moldavii. Kishinev: Shtiinitca, 1967, pp. 146–156.
276.
MlíkovskýJ. Tertiary avian localities of Russia (European part). In: MlíkovskýJ (ed) Tertiary Avian Localities of Europe. 1996, pp. 711–713.
277.
Burchak-AbramovichN. Fossil of ostrich egg shell (Struthio sp.) in Azerbaijan. Izv Akad Nauk Azerbaydzhanskoy SSR1952; 1: 39.
278.
HarrisJMLeakeyMG. Lothagam birds. In: LeakeyMGHarrisJM (eds) Lothagam: the Dawn of Humanity in Eastern Africa. New York: Columbia University Press, 2003, pp. 161–166.
279.
ZelenkovNV. Ardea sytchevskayae sp. nov., a new heron species (Aves: Ardeidae) from the middle Miocene of Mongolia. Paleontol J2011; 45: 572–579.
280.
PickfordMAl-KindiMRajhiM, et al.Correlation of the Marsawdad Formation, Oman, Late Miocene (Turolian-Ventian), based on fossil avian eggshells. Estud Geológicos2023; 79: e153.
281.
RobinsonPWimanS.K.A revision of the stratigraphic subdivision of the Miocene rocks of Sub-Dorsale Tunisia. Notes Serv Géologique Tunis1976; 70–86.
282.
WangSHuYWangL. New ratite eggshell material from the Miocene of Inner Mongolia, China. Avian Res2023; 2: 18–26.
283.
AnderssonJG. An early Chinese culture. China: Ministry of Agriculture and Commerce / Geological Survey of China, 1923.
284.
KurochkinYNLunguAN. A new ostrich from the middle Sarmatian of Moldavia. Paleont J1970; 4: 103–111.
285.
Burchak-AbramovichNVekuaAK. The Fossil Ostrich from the Akchagil lsyers of Georgia. Acta Zool Cracoviensia1971; 16: 1–28.
286.
AmoOLaplanaCLiñánE. El primer huevo fósil del Terciario aragonés. Nat Aragon2000; 5: 19–20.
287.
SauerRotheSauerFEG, et al.Ratite eggshells from Lanzarote, Canary Islands. Science1972; 176: 43–45.
288.
SánchezMarco A. New data and an overview of the past avifaunas from the Canary Islands. Ardeola2010; 57: 13–40.
289.
MlíkovskýJ. Tertiary avian localities of Ukraine. In: MlíkovskýJ (ed) Tertiary Avian Localities of Europe. 1996, pp. 743–757.
290.
SauerEGFSauerEM. Ratite eggshell fragments from Mio-Pliocene-continental sediments in the district of Ourzazate, Morocco. Palaeontogr Abt Palaeozoologie-Stratigr1978; 161: 1–54.
291.
WetmoreA. A Pleistocene Egg from Nevada. Condor1939; 41: 98–99.
292.
WilliamsDLGVickers-RichP. Fossil eggs from the Tertiary and Quaternary of Australia. Vertebr Palaeontol Australas1991; 871–891.
293.
Grellet-TinnerGSpoonerNWorthyTH. Is the “Genyornis” egg of a mihirung or another extinct bird from the Australian dreamtime?Quat Sci Rev2016; 133: 147–164.
294.
Cruz-ElizaldeRRamírez-BautistaALozanoA, et al.Reproductive strategies of a lizard community from an arid environment of Mexico. J Arid Environ2023; 208: 104875.
295.
LewisFP. Fossil eggs in Bermuda. Hobbies1928; 8: 19–23.
296.
YoungCC. New material of ostrich eggs and its stratigraphic significance. Paleovertebrata Paleoanthropologia1960; 2: 115–119.
297.
YoungCC. On the New Finds of Fossil Eggs of Struthio Anderssoni Lowe in North China with Remarks on the Egg Remains Found in Shansi, Shensi and in Choukoutien§. Bull Geol Soc China1933; 12: 145–152.
298.
MohabeyDM. Avian Egg Shells from Pleistocene of Kutch. J Geol Soc India1989; 10: 136–202.
299.
GillBT. Morphometrics of Moa eggshells fragments (Aves:Dinornithformes) from Late Holocene dune-sandsof theKarikari Peninsula, New Zealand. J R Soc N Z2000; 30: 131–145.
300.
Corona MartínezE. Las aves del Cenozoico tardío de México. Un análisis paleobiológico. Servicio de Publicaciones de la Universidad Autónoma de Madrid, 2009.
301.
CoronaMartínez M. The Pleistocene bird record of México. Acta Zool Cracoviensia2002; 45: 293–306.
302.
EmslieSDPattersonWP. Abrupt recent shift in δ13 C and δ15 N values in Adélie penguin eggshell in Antarctica. Proc Natl Acad Sci U S A2007; 104: 11666–11669.
303.
TorcelliAJ. Obras completas y correspondencia científica de Florentino Ameghino. La Plata: Taller de Impresiones Oficiales del Gobierno de la Provincia de Buenos Aires.
JanzLElstonRGBurrGS. Dating North Asian surface assemblages with ostrich eggshell: implications for palaeoecology and extirpation. J Archaeol Sci;36(9):1982-9. Epub ahead of print September 2009. DOI: 10.1016/j.jas.2009.05.012.
