The summer rainfall zone (SRZ) in the South African interior experienced pronounced hydrological and vegetation changes during the Holocene inferred to be driven mainly by shifts in atmospheric and oceanic circulations systems. The exact mechanisms controlling these changes are still debated. To gain better insights into the Holocene environmental changes in the South African SRZ and their driving factors, we analysed compound-specific carbon and hydrogen isotopes of plant wax n-alkanes (δ13Cwax and δDwax) from a marine sediment core covering the last 9900 years. The core has been recovered offshore the mouth of the Orange River, predominantly draining the South African summer rainfall region. Our data indicate a dry early Holocene and a gradual increase of wetter conditions with a higher abundance of C4 vegetation towards the middle Holocene. Wettest conditions occurred around 3900 cal. yr BP. The last 3900 years were characterised by a gradual aridification overlain by variable wetter conditions. During the ‘Little Ice Age’ (LIA: ca. 640–310 cal. yr BP), relatively dry conditions with elevated C4 plant contributions occurred. This opposite behaviour, that is, more C4 plant contribution during drier conditions compared to the remainder of the Holocene, points towards an influence of winter rainfall in the lower Orange River catchment during the late-Holocene and a decline in summer rainfall. We emphasise the importance of changes in the latitudinal insolation gradient (LIG) as a potentially important controlling mechanism for hydrologic and vegetation changes in the SRZ.
BadewienTVogtsADupontLet al. (2015) Influence of Late Pleistocene and Holocene climate on vegetation distributions in southwest Africa elucidated from sedimentary n-alkanes – Differences between 12°S and 20°S. Quaternary Science Reviews125: 160–171.
2.
BenitoGThorndycraftVRRicoMTet al. (2011) Hydrological response of a dryland ephemeral river to southern African climatic variability during the last millennium. Quaternary Research75(3): 471–482.
3.
BergerALoutreMF (1991) Insolation values for the climate of the last 10 million years. Quaternary Science Reviews10(4): 297–317.
4.
BertrandSHughenKSepúlvedaJet al. (2014) Late Holocene covariability of the southern westerlies and sea surface temperature in northern Chilean Patagonia. Quaternary Science Reviews105: 195–208.
5.
BirchGF (1977) Surficial sediments on the continental margin off the west coast of South Africa. Marine Geology23(4): 305–337.
6.
BirchGFDayRWDu PlessisA (1991) Nearshore Quaternary sediments on the west coast of southern Africa. Bulletin: Geological Survey of South Africa101: 14.
7.
BlameyRReasonC (2007) Relationships between Antarctic sea-ice and South African winter rainfall. Climate Research33(2): 183–193.
8.
BosmansJHCDrijfhoutSSTuenterEet al. (2012) Monsoonal response to mid-holocene orbital forcing in a high resolution GCM. Climate of the Past8(2): 723–740.
9.
BosmansJHCHilgenFJTuenterEet al. (2015) Obliquity forcing of low-latitude climate. Climate of the Past11: 1335–1346.
10.
BousmanCB (1991) Holocene paleoecology and Later Stone Age hunter-gatherer adaptations in the South African interior plateau. Ph.D. Dissertation, Southern Methodist University.
11.
BowenGJRevenaughJ (2003) Interpolating the isotopic composition of modern meteoric precipitation. Water Resources Research39(10): 1299.
12.
BremanEGillsonLWillisK (2012) How fire and climate shaped grass-dominated vegetation and forest mosaics in northern South Africa during past millennia. The Holocene22(12): 1427–1439.
13.
BrookGAScottLRailsbackLBet al. (2010) A 35ka pollen and isotope record of environmental change along the southern margin of the Kalahari from a stalagmite and animal dung deposits in Wonderwerk Cave, South Africa. Journal of Arid Environments74: 870–884.
14.
BurroughSLThomasDSG (2013) Central southern Africa at the time of the African Humid Period: A new analysis of Holocene palaeoenvironmental and palaeoclimate data. Quaternary Science Reviews80: 29–46.
15.
CampbellBMMollEJ (1977) The Forest Communities of Table Mountain, South Africa. Vegetatio34(2): 105–115.
16.
ChaseBMMeadowsME (2007) Late Quaternary dynamics of southern Africa’s winter rainfall zone. Earth-Science Reviews84(3–4): 103–138.
17.
ChaseBMThomasDSG (2006) Late Quaternary dune accumulation along the western margin of South Africa: Distinguishing forcing mechanisms through the analysis of migratory dune forms. Earth and Planetary Science Letters251(3–4): 318–333.
18.
ChaseBMThomasDSG (2007) Multiphase late Quaternary aeolian sediment accumulation in western South Africa: Timing and relationship to palaeoclimatic changes inferred from the marine record. Quaternary International166(1): 29–41.
19.
ChaseBMBoomACarrASet al. (2015a) Evolving southwest African response to abrupt deglacial North Atlantic climate change events. Quaternary Science Reviews121: 132–136.
20.
ChaseBMLimSChevalierMet al. (2015b) Influence of tropical easterlies in southern Africa’s winter rainfall zone during the Holocene. Quaternary Science Reviews107: 138–148.
21.
ChenFJiaJChenJet al. (2016) A persistent Holocene wetting trend in arid central Asia, with wettest conditions in the late-Holocene, revealed by multi-proxy analyses of loess-paleosol sequences in Xinjiang, China. Quaternary Science Reviews146: 134–146.
22.
ChevalierMChaseBM (2015) Southeast African records reveal a coherent shift from high- to low-latitude forcing mechanisms along the east African margin across last glacial–interglacial transition. Quaternary Science Reviews125: 117–130.
23.
ChevalierMChaseBM (2016) Determining the drivers of long-term aridity variability: A southern African case study. Journal of Quaternary Science31(2): 143–151.
24.
ChevalierMBrewerSChaseBM (2017) Qualitative assessment of PMIP3 rainfall simulations across the eastern African monsoon domains during the mid-Holocene and the Last Glacial Maximum. Quaternary Science Reviews156: 107–120.
25.
CockcroftMJWilkinsonMJTysonPD (1987) The application of a present-day climatic model to the late quaternary in southern Africa. Climatic Change10(2): 161–181.
26.
CollinsJASchefußEGovinAet al. (2014) Insolation and glacial–interglacial control on southwestern African hydroclimate over the past 140 000 years. Earth and Planetary Science Letters398: 1–10.
27.
ComptonJSHerbertCTHoffmanMTet al. (2010) A tenfold increase in the Orange River mean Holocene mud flux: Implications for soil erosion in South Africa. The Holocene20(1): 115–122.
28.
CookERPalmerJGD’ArrigoRD (2002) Evidence for a ‘Medieval Warm Period’ in a 1,100 year tree-ring reconstruction of past austral summer temperatures in New Zealand. Geophysical Research Letters29(14): 121–124.
29.
CordovaCE (2013) C3 Poaceae and Restionaceae phytoliths as potential proxies for reconstructing winter rainfall in South Africa. Quaternary International287: 121–140.
30.
CowlingRMEslerKJRundelPW (1999) Namaqualand, South Africa – An overview of a unique winter-rainfall desert ecosystem. Plant Ecology142(1–2): 3–21.
31.
CowlingRMRichardsonDMPierceSM (1997) Vegetation of Southern Africa. Cambridge: Cambridge University Press.
32.
DallmeyerAClaussenMFischerNet al. (2015) The evolution of sub-monsoon systems in the Afro-Asian monsoon region during the Holocene – comparison of different transient climate model simulations. Climate of the Past11(2): 305–326.
33.
DallmeyerAClaussenMWangYet al. (2013) Spatial variability of Holocene changes in the annual precipitation pattern: A model-data synthesis for the Asian monsoon region. Climate Dynamics40(11–12): 2919–2936.
34.
DansgaardW (1964) Stable isotopes in precipitation. Tellus16(4): 436–468.
35.
DavisBASBrewerS (2009) Orbital forcing and role of the latitudinal insolation/temperature gradient. Climate Dynamics32(2–3): 143–165.
36.
DavisBASBrewerS (2011) A unified approach to orbital, solar, and lunar forcing based on the Earth’s latitudinal insolation/temperature gradient. Quaternary Science Reviews30(15): 1861–1874.
37.
DenisDCrostaXBarbaraLet al. (2010) Sea ice and wind variability during the Holocene in East Antarctica: Insight on middle–high latitude coupling. Quaternary Science Reviews29(27): 3709–3719.
EtourneauJCollinsLGWillmottVet al. (2013) Holocene climate variations in the western Antarctic Peninsula: Evidence for sea ice extent predominantly controlled by changes in insolation and ENSO variability. Climate of the Past9(4): 1431–1446.
40.
FeakinsSJSessionsAL (2010) Controls on the D/H ratios of plant leaf waxes in an arid ecosystem. Geochimica et Cosmochimica Acta74(7): 2128–2141.
41.
FinchJMHillTR (2008) A late Quaternary pollen sequence from Mfabeni Peatland, South Africa: Reconstructing forest history in Maputaland. Quaternary Research70(3): 442–450.
42.
GamarraBSachseDKahmenA (2016) Effects of leaf water evaporative 2H-enrichment and biosynthetic fractionation on leaf wax n-alkane δ2H values in C3 and C4 grasses. Plant, Cell & Environment39: 2390–2403.
43.
GaoLEdwardsEJZengYet al. (2014) Major evolutionary trends in hydrogen isotope fractionation of vascular plant leaf waxes. PLoS ONE9(11): e112610.
44.
GarcinYSchwabVFGleixnerGet al. (2012) Hydrogen isotope ratios of lacustrine sedimentary n-alkanes as proxies of tropical African hydrology: Insights from a calibration transect across Cameroon. Geochimica et Cosmochimica Acta79: 106–126.
45.
GasseFChaliéFVincensAet al. (2008) Climatic patterns in equatorial and southern Africa from 30,000 to 10,000 years ago reconstructed from terrestrial and near-shore proxy data. Quaternary Science Reviews27(25–26): 2316–2340.
46.
GrayCED (2009) Characterising the Namaqualand mudbelt of southern Africa: Chronology, palynology and palaeoenvironments. South African Geographical Journal82: 137–142.
47.
GrayCEDMeadowsMELee-ThorpJAet al. (2000) Characterising the Namaqualand mudbelt of southern Africa: Chronlogy, palynology and palaeoenvironments. South African Geographical Journal82(3): 137–142.
48.
HahnAComptonJSMeyer-JacobCet al. (2016) Holocene paleo-climatic record from the South African Namaqualand mudbelt: A source to sink approach. Quaternary International404: 121–135.
49.
HeineKVölkelJ (2011) Extreme floods around AD 1700 in the northern Namib Desert, Namibia, and in the Orange River catchment, South Africa – Were they forced by a decrease of solar irradiance during the Little Ice Age?Geographia Polonia84(1): 61–80.
50.
HerbertCTComptonJS (2007) Geochronology of Holocene sediments on the western margin of South Africa. South African Journal of Geology110(2–3): 327–338.
51.
HerrmannNBoomACarrASet al. (2016) Sources, transport and deposition of terrestrial organic material: A case study from southwestern Africa. Quaternary Science Reviews149: 215–229.
52.
HerrmannNBoomACarrASet al. (2017) Hydrogen isotope fractionation of leaf wax n-alkanes in southern African soils. Organic Geochemistry109: 1–13.
53.
HijmansRJCameronSEParraJLet al. (2005) Very high resolution interpolated climate surfaces for global land areas. International Journal of Climatology25(15): 1965–1978.
54.
HodellDAKanfoushSLShemeshAet al. (2001) Abrupt cooling of Antarctic surface waters and sea ice expansion in the south Atlantic sector of the Southern Ocean at 5000 cal yr B.P. Quaternary Research56(2): 191–198.
55.
HolmgrenKLee-ThorpJACooperGRJet al. (2003) Persistent millennial-scale climatic variability over the past 25,000 years in Southern Africa. Quaternary Science Reviews22(21): 2311–2326.
56.
HouJDAndreaWJMacDonaldDet al. (2007) Hydrogen isotopic variability in leaf waxes among terrestrial and aquatic plants around Blood Pond, Massachusetts (USA). Organic Geochemistry38(6): 977–984.
57.
JiangDTianZLangX (2015) Mid-Holocene global monsoon area and precipitation from PMIP simulations. Climate Dynamics44(9–10): 2493–2512.
58.
JiangHBjörckSSvenssonN-O (1998) Reconstruction of Holocene sea-surface salinity in the Skagerrak–Kattegat: A climatic and environmental record of Scandinavia. Journal of Quaternary Science13(2): 107–114.
59.
KahmenASchefußESachseD (2013b) Leaf water deuterium enrichment shapes leaf wax n-alkane δD values of angiosperm plants I: Experimental evidence and mechanistic insights. Geochimica et Cosmochimica Acta111: 39–49.
60.
KahmenAHoffmannBSchefußEet al. (2013a) Leaf water deuterium enrichment shapes leaf wax n-alkane δD values of angiosperm plants II: Observational evidence and global implications. Geochimica et Cosmochimica Acta111: 50–63.
61.
KeelingCD (1979) The Suess effect: 13Carbon-14Carbon interrelations. Environment International2(4–6): 229–300.
62.
KristenIWilkesHViethAet al. (2010) Biomarker and stable carbon isotope analyses of sedimentary organic matter from Lake Tswaing: Evidence for deglacial wetness and early Holocene drought from South Africa. Journal of Paleolimnology44(1): 143–160.
63.
KrullESachseDMüglerIet al. (2006) Compound-specific δ13C and δ2H analyses of plant and soil organic matter: A preliminary assessment of the effects of vegetation change on ecosystem hydrology. Soil Biology and Biochemistry38(11): 3211–3221.
64.
KuechlerRRDupontLMSchefußE (2018) Hybrid insolation forcing of Pliocene monsoon dynamics in West Africa. Climate of the Past14(1): 73–84.
65.
LamyFHebbelnDRöhlUet al. (2001) Holocene rainfall variability in southern Chile: A marine record of latitudinal shifts of the Southern Westerlies. Earth and Planetary Science Letters185(3): 369–382.
66.
LeanJRindD (1999) Evaluating sun–climate relationships since the Little Ice Age. Journal of Atmospheric and Solar-Terrestrial Physics61(1–2): 25–36.
67.
LeducGHerbertCTBlanzTet al. (2010) Contrasting evolution of sea surface temperature in the Benguela upwelling system under natural and anthropogenic climate forcings. Geophysical Research Letters37(20): L20705.
68.
LiFWuN (2010) Pliocene land snail record from western Chinese Loess Plateau and implications for impacts of the summer insolation gradient between middle and low latitudes on the East Asian summer monsoon. Global and Planetary Change72(1): 73–78.
69.
LimSChaseBMChevalierMet al. (2016) 50,000 years of vegetation and climate change in the Southern Namib Desert, Pella, South Africa. Palaeogeography, Palaeoclimatology, Palaeoecology451: 197–209.
70.
LiuWYangH (2008) Multiple controls for the variability of hydrogen isotopic compositions in higher plant n-alkanes from modern ecosystems. Global Change Biology14(9): 2166–2177.
71.
LiuWYangHLiL (2006) Hydrogen isotopic compositions of n-alkanes from terrestrial plants correlate with their ecological life forms. Oecologia150(2): 330–338.
72.
LodderJ (2011) The late quaternary palaeoenvironments of a subalpine wetland in Cathedral Peak, KwaZulu-Natal Drakensberg. Master’s Thesis, University of Kwazulu-Natal.
73.
LoutreM-FPaillardDVimeuxFet al. (2004) Does mean annual insolation have the potential to change the climate?Earth and Planetary Science Letters221(1): 1–14.
74.
LuJVecchiGAReichlerT (2007) Expansion of the Hadley cell under global warming. Geophysical Research Letters34(6): L06805.
75.
LuoPPengPGleixnerGet al. (2011) Empirical relationship between leaf wax n-alkane δD and altitude in the Wuyi, Shennongjia and Tianshan Mountains, China: Implications for paleoaltimetry. Earth and Planetary Science Letters301(1–2): 285–296.
76.
LyonsRToothSDullerGAT (2014) Late Quaternary climatic changes revealed by luminescence dating, mineral magnetism and diffuse reflectance spectroscopy of river terrace palaeosols: A new form of geoproxy data for the southern African interior. Quaternary Science Reviews95: 43–59.
77.
MaboteMERogersJMeadowsME (1997) Sedimentology of terrigenous mud from the Orange River delta and the inner shelf off Namaqualand, South Africa. South African Geographical Journal79(2): 108–114.
78.
McInerneyFAHellikerBRFreemanKH (2011) Hydrogen isotope ratios of leaf wax n-alkanes in grasses are insensitive to transpiration. Geochimica et Cosmochimica Acta75(2): 541–554.
79.
Masson-DelmotteVJouzelJLandaisAet al. (2005) GRIP deuterium excess reveals rapid and orbital-scale changes in Greenland moisture origin. Science309(5731): 118–121.
80.
MeadowsMERogersJLee-ThorpJAet al. (2002) Holocene geochronology of a continentalshelf mudbelt off southwestern Africa. The Holocene12: 59–67.
81.
MetwallyAAScottLNeumannFHet al. (2014) Holocene palynology and palaeoenvironments in the Savanna Biome at Tswaing Crater, central South Africa. Palaeogeography, Palaeoclimatology, Palaeoecology402: 125–135.
82.
MooneyHATroughtonJHBerryJA (1977) Carbon isotope ratio measurements of succulent plants in southern Africa. Oecologia30(4): 295–305.
83.
MoossenHBendleJSekiOet al. (2015) North Atlantic Holocene climate evolution recorded by high-resolution terrestrial and marine biomarker records. Quaternary Science Reviews129: 111–127.
84.
MucinaLRutherfordM (2006) The vegetation of South Africa, Lesotho and Swaziland. Pretoria: South African National Biodiversity Institute, pp. 749–790.
85.
NashDJDe CortGChaseBMet al. (2016) African hydroclimatic variability during the last 2000 years. Quaternary Science Reviews154: 1–22.
86.
NeumannFHBothaGAScottL (2014) 18,000 years of grassland evolution in the summer rainfall region of South Africa: Evidence from Mahwaqa Mountain, KwaZulu-Natal. Vegetation History and Archaeobotany23(6): 665–681.
87.
NeumannFHScottLBousmanCBet al. (2010) A Holocene sequence of vegetation change at Lake Eteza, coastal KwaZulu-Natal, South Africa. Review of Palaeobotany and Palynology162(1): 39–53.
88.
NorströmENeumannFHScottLet al. (2014) Late Quaternary vegetation dynamics and hydro-climate in the Drakensberg, South Africa. Quaternary Science Reviews105: 48–65.
89.
NorströmEScottLPartridgeTCet al. (2009) Reconstruction of environmental and climate changes at Braamhoek wetland, eastern escarpment South Africa, during the last 16,000 years with emphasis on the Pleistocene–Holocene transition. Palaeogeography, Palaeoclimatology, Palaeoecology271(3–4): 240–258.
90.
PAGES 2k Consortium (2013) Continental-scale temperature variability during the past two millennia. Nature Geoscience6: 339–346.
91.
PartridgeTC (1997) Cainozoic environmental change in southern Africa, with special emphasis on the last 200 000 years. Progress in Physical Geography21(1): 3–22.
92.
PartridgeTCDemenocalPBLorentzSAet al. (1997) Orbital forcing of climate over South Africa: A 200,000-year rainfall record from the pretoria saltpan. Quaternary Science Reviews16(10): 1125–1133.
93.
PolissarPJFreemanKH (2010) Effects of aridity and vegetation on plant-wax δD in modern lake sediments. Geochimica et Cosmochimica Acta74(20): 5785–5797.
94.
RamischALockotGHaberzettlTet al. (2016) A persistent northern boundary of Indian Summer Monsoon precipitation over Central Asia during the Holocene. Scientific Reports6: 25791.
95.
ReasonCJCRouaultM (2002) ENSO-like decadal variability and South African rainfall. Geophysical Research Letters29(13): 1638.
96.
ReasonCJCRouaultM (2005) Links between the Antarctic Oscillation and winter rainfall over western South Africa. Geophysical Research Letters32(7): L07705.
97.
ReasonCJCRouaultMMeliceJ-Let al. (2002) Interannual winter rainfall variability in SW South Africa and large scale ocean-atmosphere interactions. Meteorology and Atmospheric Physics80(1–4): 19–29.
98.
RepinskiPHolmgrenKLauritzenSEet al. (1999) A late Holocene climate record from a stalagmite, Cold Air Cave, Northern Province, South Africa. Palaeogeography, Palaeoclimatology, Palaeoecology150(3): 269–277.
99.
RichardsonDM (2000) Ecology and Biogeography of Pinus. Cambridge: Cambridge University Press.
100.
RisebrobakkenBDokkenTOtteråOHet al. (2007) Inception of the Northern European ice sheet due to contrasting ocean and insolation forcing. Quaternary Research67(1): 128–135.
101.
RogersJBremnerJM (1991) The Benguela ecosystem. Part VII. Marine-geological aspects. Oceanography and Marine Biology: An Annual Review29: 1–85.
102.
RogersJRauA (2006) Surficial sediments of the wave-dominated Orange River Delta and the adjacent continental margin off south-western Africa. African Journal of Marine Science28(3–4): 511–524.
103.
SachseDRadkeJGleixnerG (2004) Hydrogen isotope ratios of recent lacustrine sedimentary n-alkanes record modern climate variability. Geochimica et Cosmochimica Acta68(23): 4877–4889.
104.
SchaeferJMDentonGHKaplanMet al. (2009) High-frequency Holocene glacier fluctuations in New Zealand differ from the Northern Signature. Science324(5927): 622–625.
105.
SchneiderRBleilUBuschhoffHet al. (2003) Report and preliminary results of Meteor cruise M57/1, Cape Town - Walvis Bay, 20.01. - 08.02.2003. No. 216.Bremen: Berichte aus dem Fachbereich Geowissenschaften der Universität Bremen.
106.
SchwabVFGarcinYSachseDet al. (2015) Effect of aridity on δ13C and δD values of C3 plant- and C4 graminoid-derived leaf wax lipids from soils along an environmental gradient in Cameroon (Western Central Africa). Organic Geochemistry78: 99–109.
107.
ScottLNyakaleM (2002) Pollen indications of Holocene palaeoenvironments at Florisbad spring in the central Free State, South Africa. The Holocene12(4): 497–503.
108.
ScottLVogelJC (2000) Evidence for environmental conditions during the last 20000 years in Southern Africa from 13C in fossil hyrax dung. Global and Planetary Change26(1): 207–215.
109.
ScottLWoodborneS (2007) Vegetation history inferred from pollen in Late Quaternary faecal deposits (hyraceum) in the Cape winter-rain region and its bearing on past climates in South Africa. Quaternary Science Reviews26: 941–953.
110.
ScottLBousmanCBNyakaleM (2005) Holocene pollen from swamp, cave and hyrax dung deposits at Blydefontein (Kikvorsberge), Karoo, South Africa. Quaternary International129(1): 49–59.
111.
ScottLNeumannFHBrookGAet al. (2012) Terrestrial fossil-pollen evidence of climate change during the last 26 thousand years in Southern Africa. Quaternary Science Reviews32: 100–118.
112.
ShiNDupontLMBeugH-Jet al. (2000) Correlation between vegetation in southwestern Africa and oceanic upwelling in the past 21,000 years. Quaternary Research54(1): 72–80.
113.
SingarayerJSBurroughSL (2015) Interhemispheric dynamics of the African rainbelt during the late Quaternary. Quaternary Science Reviews124: 48–67.
114.
SmithFAFreemanKH (2006) Influence of physiology and climate on δD of leaf wax n-alkanes from C3 and C4 grasses. Geochimica et Cosmochimica Acta70(5): 1172–1187.
115.
StagerJCMayewskiPAWhiteJet al. (2012) Precipitation variability in the winter rainfall zone of South Africa during the last 1400 yr linked to the austral westerlies. Climate of the past8(3): 877–887.
116.
StrelinJCasassaGRosqvistGet al. (2008) Holocene glaciations in the Ema Glacier valley, Monte Sarmiento Massif, Tierra del Fuego. Palaeogeography, Palaeoclimatology, Palaeoecology260(3): 299–314.
117.
StuutJ-BWCrostaXvan der BorgKet al. (2004) Relationship between Antarctic sea ice and southwest African climate during the late Quaternary. Geology32(10): 909.
118.
SuessHE (1955) Radiocarbon concentration in modern wood. Science122(3166): 415–417.
119.
TalmaASVogelJC (1992) Late Quaternary paleotemperatures derived from a speleothem from Cango Caves, Cape Province, South Africa. Quaternary Research37(2): 203–213.
120.
TaylorA (2004) A trace element study of sediments from the Olifants River estuary, the Berg River estuary, and the off-shore Mud Belt. Available at: https://open.uct.ac.za/handle/11427/9027
121.
ThackerayJFLee-ThorpJA (1992) Isotopic analysis of equid teeth from Wonderwerk Cave, northern Cape Province, South Africa. Palaeogeography, Palaeoclimatology, Palaeoecology99(1): 141–150.
122.
ThevenonFWilliamsonDTaiebM (2002) A 22 kyr BP sedimentological record of Lake Rukwa (8°S, SW Tanzania): Environmental, chronostratigraphic and climatic implications. Palaeogeography, Palaeoclimatology, Palaeoecology187(3): 285–294.
123.
TrucLChevalierMFavierCet al. (2013) Quantification of climate change for the last 20,000years from Wonderkrater, South Africa: Implications for the long-term dynamics of the Intertropical Convergence Zone. Palaeogeography, Palaeoclimatology, Palaeoecology386: 575–587.
124.
TysonPDPreston-WhyteRA (2000) The Weather and Climate of Southern Africa. Cape Town: Oxford University Press.
125.
ValsecchiVChaseBMSlingsbyJAet al. (2013) A high resolution 15,600-year pollen and microcharcoal record from the Cederberg Mountains, South Africa. Palaeogeography, Palaeoclimatology, Palaeoecology387: 6–16.
126.
VerschurenDSinninghe DamstéJSMoernautJet al. (2009) Half-precessional dynamics of monsoon rainfall near the East African Equator. Nature462(7273): 637–641.
127.
VimeuxFMassonVJouzelJet al. (2001) Holocene hydrological cycle changes in the Southern Hemisphere documented in East Antarctic deuterium excess records. Climate Dynamics17(7): 503–513.
128.
VoarintsoaNRGBrookGALiangFet al. (2016) Stalagmite multi-proxy evidence of wet and dry intervals in northeastern Namibia: Linkage to latitudinal shifts of the Inter-Tropical Convergence Zone and changing solar activity from AD 1400 to 1950. The Holocene27: 384–396.
129.
VogelJCFulsAEllisRP (1978) The geographical distribution of Kranz grasses in South Africa. South African Journal of Science74: 209–215.
WannerHBeerJBütikoferJet al. (2008) Mid- to Late Holocene climate change: An overview. Quaternary Science Reviews27(19–20): 1791–1828.
132.
WeldeabSLeaDWOberhänsliHet al. (2014) Links between southwestern tropical Indian Ocean SST and precipitation over Southeastern Africa over the last 17 kyr. Palaeogeography, Palaeoclimatology, Palaeoecology410: 200–212.
133.
WeldeabSStuutJ-BWSchneiderRRet al. (2013) Holocene climate variability in the winter rainfall zone of South Africa. Climate of the past Discussions9(3): 2309–2356.
134.
WergerMJAEllisRP (1981) Photosynthetic pathways in the arid regions of South Africa. Flora171: 64–75.
135.
YamoriWHikosakaKWayDA (2014) Temperature response of photosynthesis in C3, C4, and CAM plants: Temperature acclimation and temperature adaptation. Photosynthesis Research119(1–2): 101–117.
136.
YoungMABradleyRS (1984) Insolation gradients and the paleoclimatic record. In: BergerAImbrieJHaysJet al (eds) Milankovitch and Climate. Dordrecht: Springer, pp. 707–713.
137.
ZawadaPKHattinghJVan BladerenDet al. (1996) Paleoflood Hydrological Analysis of Selected South African Rivers. Pretoria: Water Research Commission.
138.
ZhangQHolmgrenKSundqvistH (2015) Decadal rainfall dipole oscillation over southern Africa modulated by variation of austral summer land–sea contrast along the east coast of Africa. Journal of the Atmospheric Sciences72(5): 1827–1836.
139.
ZhaoXDupontLMeadowsMEet al. (2015) Pollen distribution in the marine surface sediments of the mudbelt along the west coast of South Africa. Quaternary International404: 44–56.
140.
ZhaoXDupontLSchefußEet al. (2017) Palynological evidence for Holocene climatic and oceanographic changes off western South Africa. Quaternary Science Reviews165: 88–101.
141.
ZhaoXDupontLSchefußEet al. (2016) Holocene vegetation and climate variability in the winter and summer rainfall zones of South Africa. The Holocene26: 843–857.
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