Abstract
This article considers the sexual politics of animal evidence in the context of German sexology around 1920. In the 1910s, the German-Jewish geneticist Richard B. Goldschmidt conducted experiments on the moth
In 1930, Magnus Hirschfeld published the fourth volume of his

The butterfly in question was named after Hirschfeld by the Austrian-Swedish entomologist and anthropologist Felix Bryk in a 1929 issue of the German entomology journal
Over a decade before
In ‘“Unnatural Acts” in Nature: The Scientific Fascination With Queer Animals’, Jennifer Terry examines the ways in which geneticists, primatologists, biologists, and popular culture observe same-sex behaviour in animals. She asks, ‘What possibilities do animals, behaving in a “queer” manner, open up for humans interested in making sense of sexuality?’ (Terry, 2000: 152). This question is central to my work on the role of animals in the development of sexological debate, too, although the emphasis in sexological responses to Goldschmidt’s butterfly research is less on behaviour than on discussions of homosexual constitution, a sense of
To put sexologists’ interest in intersex butterflies in context, Goldschmidt’s butterflies/moths were not the only example of the political authority of animal nature in interwar sexology. The interconnected web binding together sex, development, and hormones featured prominently in the work of the Austrian endocrinologist Eugen Steinach. Steinach dedicated himself to the importance of the inner secretions (since 1905 also referred to as hormones) for the study of sex. He hypothesised that ‘the characteristics of sex could always be modified by modulating the functions of the sex glands’ (Sengoopta, 2006: 65). Steinach’s experimental work with guinea pigs and rats focused on same-sex and cross-sex transplants of gonads and concluded that the inner secretions produced masculine and feminine characteristics, arguing that sexuality was flexible and changeable rather than given in the gametes (Logan, 2013: 30–3). Hirschfeld took notice of Steinach’s work, and adopted his glandular theory of homosexuality to support his biological model of homosexuality as non-pathological and organic (Sengoopta, 1998). Hirschfeld showed a great interest in Steinach’s animal experiments, arguing that ‘nature had already produced human beings with the features of Steinach’s feminized and masculinized animals’ (ibid.: 464). As Sengoopta argues, Hirschfeld here used Steinach (and his experimental animals) as political allies for his sexual rights programme (ibid.). Steinach in turn drew on Hirschfeld’s clinical work to argue that lacking gonadal differentiation caused homosexuality. Convinced by Hirschfeld that his experimental work could be applied to humans, Steinach ‘turned enthusiastically from guinea pigs to men’ (ibid.: 465), and collaborated with the urologist Robert Lichtenstern on gonadal grafting in humans (Oudshoorn, 1994: 57). Hirschfeld and his colleagues adopted this practice. For Hirschfeld, Steinach’s animal experiments confirmed that homosexuality was indeed of a biological nature, with direct political implications: homosexuality was part of a natural spectrum of sexuality and could not be criminalised.
Sexologists evidently had an interest in animal evidence more broadly, but it is important to keep in mind the specificity of the particular animal used as a form of evidence for human sexuality. Whereas some geneticists, such as Thomas Hunt Morgan, built their experimental work on
In a post-Darwinian context, the marshalling of animal evidence by sexologists to explain human conditions is not surprising, but what kind of evidence is used, the framework of thought implied, and to what ends it is used can have political implications. In
It is this intellectual work that I want to examine in this article: How was knowledge about butterflies transferred to humans and to what ends? What scientific and methodological frameworks were used to facilitate this transmission? One aspect of this intellectual work was the transmission of evidence across disciplinary lines: Using the experimental work on intersex butterflies in the context of sexological work required a translation from one disciplinary context to another, from zoology and animal genetics to sexology. By paying attention to this interdisciplinary exchange between animal genetics and sexology, this article also builds on recent work on the interdisciplinary nature of sexology (Fisher and Funke, 2015, 2018). Tracing this cross-disciplinary and often interdisciplinary exchange of evidence shows that the meaning and significance of butterfly evidence was not fixed, as evidence had to be fitted into an already existing but continually developing set of sexological hypotheses and methodologies. The circulation of these claims amongst sexologists put them into a different sociopolitical context, with ramifications for how they could be understood both scientifically and politically. The necessity of marshalling butterfly evidence lay in the fact that particular authority was accorded to natural and biological science. By relying on evidence from animal genetics, sexologists thereby also used this cross-disciplinary knowledge transfer to support sexological theories and widen their field of influence. In the following article, then, I explore the complex politics of evidence that took place when intersex butterflies were used to inform central sexological questions, most importantly the debate around the nature and naturalness of sexual intermediacy. 3
Sex and race in intersex butterflies: Richard B. Goldschmidt
German-Jewish geneticist Richard Benedict Goldschmidt pioneered research on intersex butterflies. Goldschmidt studied medicine and zoology, and in 1914 was appointed as the head of his own department, focusing on animal genetics and biology, at the new Kaiser Wilhelm Institute (KWI) for Biology in Dahlem, Berlin (Goldschmidt, 1960: 79–81). Goldschmidt’s research depended on access to geographically diverse specimens of his experimental organism, the gypsy moth
Goldschmidt’s work was part of a larger scientific effort of biologists to understand the nature of sex. In
In the midst of these debates, the significance of Mendel’s laws of heredity, first developed in the 1860s, was rediscovered by biologists. Mendel’s experiments on pea plants showed that individual traits are determined by pairs of factors (alleles) that separate during gamete formation and randomly recombine, producing genetic diversity. In the early 1900s, chromosomes were suspected of being these carriers of genetic material. The ‘odd chromosomes’ discovered by Stevens and Wilson were increasingly called ‘sex chromosomes’. In the early decades of the 20th century, geneticists like Goldschmidt now set out to develop theories of the gene to investigate what aspect of the sex chromosomes determined sex from one generation to another, and whether this took place according to Mendelian laws of heredity. In the late 19th century, Richardson asserts, ‘researchers…believed that sex was a continuous (spectrum) rather than a discontinuous (binary) trait’ (Richardson, 2013: 24). By the 1930s, sex chromosomes had been drawn into the centre of molecular biology, and ‘today’s notion of the X and Y as the molecular agents of sex and as the genetic homunculi underlying sexual dimorphism—as “sex itself”—[was] first recognizable’ (ibid.: 71). But during Goldschmidt’s early years and the early years of genetics leading up to this moment, hormonal and genetic sex determination was considered complex and fluid.
Goldschmidt was an influential, prominent, but also controversial figure in the nascent field of genetics arising in this context of sex research. Michael Dietrich has written extensively about Goldschmidt’s ‘heresies’ in interwar genetics, which questioned the orthodoxy of the American evolutionary biologist Thomas Hunt Morgan (1866–1945). Whereas Morgan focused on gene transmission alone, Goldschmidt’s theory focused on quantitative and physiological aspects, and on gene action and function (Dietrich, 1995: 435). In the 1930s, his work underwent a major change as he began to argue that the chromosome, rather than the gene, was the unit of heredity (ibid.: 436). This dissenting view led to serious criticism of Goldschmidt’s work and his standing as an influential figure in genetics. Dietrich argues that Goldschmidt’s reputation as iconoclast was cemented by the response to his 1940 book
Goldschmidt’s experimental work of the early 1910s investigated the heredity of sex difference using different ‘races’ of the gypsy moth from Japan and Europe.
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‘Race’ here refers to geographically distinct populations of an animal species (Goldschmidt, 1915: 565). Certain combinations of different populations resulted in offspring that were no longer sexually dimorphic and displayed intermediate sexual characteristics (Dietrich, 2016: 26). Rather than combining male and female characteristics, however, these individuals were shown to have developed initially as female and then developed as male later on, and vice versa (Richmond, 2007: 177). As a result, Goldschmidt called these individuals ‘intersex’. Goldschmidt hypothesised that intersexuality was caused by a difference in ‘potency’ or ‘valence’. According to Goldschmidt, each organism contained the predisposition for either sex. Within each organism, a quantitative value of the sex factor –– its ‘valence’ or ‘potency’ –– and its relation to the potency of the predisposition of the other sex, decided which one finally appeared (Goldschmidt, 1916: 709). Goldschmidt argued that the quantitative values of the sex factor varied across geographical regions. He therefore used various natural populations of the gypsy moth from different geographic regions in Europe and Japan in order to study sex expressions in genitalia, antennae, colouration of the wings, and so on, all of which were particularly distinct in female and male individuals of

A series of intersexual females of
Scientific discussions of the sex problem were closely embedded in the political, social, and cultural circumstances of the early 20th century. They were informed by and informing broad debates about eugenics, feminism, and theories of race and ‘miscegenation’.
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Koehler and Satzinger show that Goldschmidt’s scientific discussion of sex determination, and the wider context of Goldschmidt’s colleagues and rivals, was profoundly tied up with the larger social concern of modernity: a changing context of gender, gender relations, and sexuality, in particular homosexuality (Koehler, 1998: ii; Satzinger, 2009b: 159). Goldschmidt’s work, which, he argued, showed that there was no clear-cut two-sex system, but that sexual difference existed on a sliding scale, was particularly pertinent in the Weimar Republic, where the social order of gender was rapidly evolving (Satzinger, 2009b). In this context, Goldschmidt’s butterfly experiments were co-opted for various political uses. Whilst this article focuses on the ways in which Goldschmidt’s butterflies were made to serve sexological arguments about human sexual diversity, his work was also incorporated into specifically right-wing and
As a consequence, sexologists’ interest in intersex butterflies has to be considered as part of the broader attempts to accommodate Goldschmidt’s work in very different models of race, gender, and sexuality. In 1916, Goldschmidt catapulted himself directly into these political discussions by publishing an article in which he argued that his butterfly experiments were of relevance to the study of human intersex, suggesting that homosexuality was one such form of intersex in humans (Goldschmidt, 1916/18). 7 Although Goldschmidt’s work hardly ever commented on issues as politically charged as the topic of homosexuality, there are several reasons why he felt it appropriate to comment on this occasion. As Koehler argues, ‘Goldschmidt…sought to produce a single universal theory of sex.…Goldschmidt used his work on intersexuality to explain human homosexuality, and argued that the phenomena of sex were fundamentally the same’ (Koehler, 1998: 57–8). Goldschmidt chose to focus on (male) homosexuality as a form of intersex as the simplest choice from a selection of more difficult options (ibid.: 221). Beyond this simple suitability of the topic of male homosexuality, Koehler argues that Goldschmidt was frustrated with the likes of Hirschfeld, who lacked specialist knowledge in the biological sciences and genetics, weighing in on the sex question (ibid.: 198). Goldschmidt wrote, ‘there is hardly another problem which has been such a playground of dilettantism, and if we look through the older literature on the sex problem, we find almost as many philosophers and economists inventing sex-theories as there are biologists’ (Goldschmidt, 1916: 705). Dietrich understands Goldschmidt’s comment here as a reference to the ‘growing number of sex researchers and the new field of sexual science’, arguing that ‘Goldschmidt used the experimental nature of his research as a means both of claiming greater authority to speak on the science of intersexuality and of differentiating himself from Hirschfeld and others’ (Dietrich, 2000: 225).
In the process of writing his article about human homosexuality from the standpoint of animal genetics, Goldschmidt familiarised himself with sexological literature and contributed to an interdisciplinary debate about the nature of homosexuality. Researching for his article on human hermaphroditism whilst in the USA, he asked the director of the Eugenics Record Office, Charles Davenport, for assistance in finding copies of the
Despite his apparent conviction that the sex question was best left to biologists and geneticists, Goldschmidt, too, tentatively contributed to debates outside his immediate field of expertise, and spoke in favour of such interdisciplinary work. He opened his article on human hermaphroditism with the words: The infinite specialization of science [
The sexual politics of scientific practice: Max Hodann
In 1917, a review article by Max Hodann (1894–1946), entitled ‘Neue Forschungen zur Kenntnis der hereditär-physiologischen Grundlagen sexueller Zwischenstufen’ (New Research on the Hereditary-Physiological Foundations of Sexual Intermediacy), was published in the
Hodann’s interest in sexology was sparked by an encounter with Hirschfeld at a talk he gave in 1915 (Herzer, 1985). The article about Goldschmidt’s intersex butterflies in the
If we accepted this, Goldschmidt argued (and Hodann agreed), several conclusions presented themselves. First, intersexuality arose from the crossing of geographically diverse individuals. Whereas Goldschmidt’s article went on to explain the complexity of this issue, arguing that human populations were always already ‘racially’ mixed – that is, geographically distinct (Goldschmidt, 1916/18: 8) – Hodann reduced this conclusion to the simple statement that ethnographic data might help the sexologist understand the phenomenon of human intersexuality (Hodann, 1917: 65). He reduced Goldschmidt’s complex argument and focused on the importance of ethnographic data for the study of sex in order to show the compatibility with Hirschfeld’s work. Hirschfeld showed a keen interest in ethnographic material (see Bauer, 2011; Fisher and Funke, 2018). Another conclusion that Hodann drew alongside Goldschmidt was that in humans, unlike insects, the inner secretions played an important role in sexual development (Goldschmidt, 1916/18: 12; Hodann, 1917: 67). Goldschmidt tentatively argued that ‘intersexuality could be “cured” through treatment with gonadal extract or transplantation of normal gonads’ (Goldschmidt, 1916/18: 14). Framing a cure in quotation marks suggests that Goldschmidt did not consider intersexuality/homosexuality a disease, and indeed the following conclusion confirmed this: An intersexual individual could not be assigned a sex, because they were not fully male or female (ibid.). Although he did not follow this with a statement about the decriminalisation of homosexuality, it is clear that Goldschmidt understood intersexuality as part of natural sexual variation. Hodann’s article also pointed out this ‘healing effect’ (‘Heilwirkung’; Hodann, 1917: 67) of extract and transplant therapy. Whereas Goldschmidt was less interested in the clinical application of his work, Hodann, whose work focused on the clinical application of sex research, took the potential for hormone and gonad treatment into serious consideration. Again, this was in line with Hirschfeld’s interest in these procedures in his search for a possible proof for the biological foundation of (homo)sexuality (Beachy, 2014: 175).
Hodann would have relied on Hirschfeld’s clinical material to inform his thinking, because, at the time of writing, he had not finished his medical degree and would not have had extensive clinical experience himself. In this way, Hodann attempted to establish both himself and sexology. In 1917, sexology was still in the process of establishing itself as a scientifically recognised field. Hirschfeld was less than two years away from founding his Institute of Sexology. Any cross-disciplinary link with the well-established and well-funded biological sciences could only help the sexological cause. In demonstrating his allegiance to Hirschfeld’s sexological theories by contributing a link with biological sciences, Hodann made a space for himself in sexological circles, both figuratively and literally speaking: In the mid-1920s, he joined the staff of the Institute of Sexology. Using Goldschmidt’s butterfly experiments from the field of animal genetics to support a hypothesis from the emerging field of sexology was therefore also a political move to support sexology’s credibility.
Hodann then asked what Physical and psychological intersexuality of all stages exists. An individual that is genetically supposed to belong to one sex is, in fact, intersex. From this it follows that –– because laws cannot be based on chromosomes, but on the actual situation of an individual –– no court of law is able to assign sex/gender…because this individual belongs, in actual fact, to neither of the two sexes. (Goldschmidt, 1916/18: 14, cited in Hodann, 1917: 68)
To this, Hodann added his own conclusion: ‘Those who have had the opportunity to observe the sheer diversity of sexual intermediate types in life, as well as in physical and psychological manifestations, can only agree with Goldschmidt’s…conclusions’ (Hodann, 1917: 68). Here, he returned to the idea that experimental work on butterflies conducted in a controlled laboratory environment and clinical observation stood in dialogue, and showed comparable results. For Hodann, the natural diversity of sexual life – in both butterflies and humans – had political consequences. Whilst Hodann was establishing himself as an ally of sexological causes, he was also particularly interested in social hygiene, public medicine, and sex education, spurred on by his political affiliations with the German socialist movement. When he joined the staff at the Institute of Sexology in the mid-1920s, he did so in the capacity of sex educator/adviser (Sigusch and Grau, 2009: 296–302). He lectured on topics including abortion, contraception, masturbation, and sexuality. He also wrote sex education material from a socialist perspective specifically for working-class children (Sauerteig, 2009: 133–5; 2012). As Atina Grossmann argues, ‘Wherever the ubiquitous Hodann published, he always used his dual authority as physician and working-class activist to plead for tolerance’ (Grossman, 1995: 29). In his use of Goldschmidt’s experimental work on butterflies, Hodann referred to the natural diversity and variation of sex, which depended on ‘potency’, rather than a clear and definite appearance of sex, to argue that the law should recede, because it was not fit to categorise people into male and female. The argument was not that we should do what nature said (he explicitly said he did not want to assign sex according to chromosomes), but that the different layers of sex added up to a complex system that could not be adequately represented. Intersex butterflies were used to make the argument that natural variation exceeded its legal comprehension. This line of argument was in line with Hodann’s general approach to the scientific study of sex, which considered medicine and science as fundamentally political.
In Hodann’s review of Goldschmidt’s work, intersex butterflies were mirror images of sexual intermediates in the human realm. This was reflected by the larger disciplinary twinning Hodann conducted of Goldschmidt’s experimental work and Hirschfeld’s clinical observations. Both experimental and clinical evidence, Hodann argued, provided a natural basis for tolerance. In this way, the intersexuality of butterflies was marshalled to provide a basis for tolerance of sexual diversity in the human realm, and thereby also clarified Hodann’s position as advocator for Hirschfeld’s work, which would later secure him a position at the Institute. In Hodann’s case, butterfly evidence was political in a double sense: It was marshalled to support a political aim, namely the recognition of sexual diversity beyond a limiting two-sex model, and it was a political move to establish the emerging field of sexology as equal to the more established biological sciences.
The science behind gay rights activism: Eugène Wilhelm
Three years after Hodann, Eugène Wilhelm (1866–1951) published another review of Goldschmidt’s article (Praetorius, 1920). Wilhelm was a lawyer and sexologist originally from France. He was not medically trained, but he had a significant influence on sexology because from 1900 to 1922 he wrote the ‘Bibliography of Homosexuality’, the review section of the
In his review of Goldschmidt’s work from 1920, Wilhelm, like Hodann, faithfully summarised Goldschmidt’s argument, but where Hodann focused on the comparability of experimental and clinical work to show the natural variation of sexual intermediacy, Wilhelm was interested exclusively in the meaning of Goldschmidt’s work for the argument that homosexuality was inborn. Wilhelm argued that homosexuality was not acquired through external factors, but was deeply linked to the body’s earliest developmental stages. Goldschmidt’s research, he argued, proved this: Homosexuality was one form of intersex, and Goldschmidt’s butterfly experiments had shown that these were linked to an individual’s genetic predisposition. Whereas Hodann saw hereditary experiments on butterflies and clinical observation of human patients as producing comparable outcomes, when Wilhelm introduced Goldschmidt’s work, he did not mention that his model organism was the gypsy moth, but simply talked about intersex in general terms: ‘As such, when two races (
For Wilhelm, Goldschmidt’s butterflies were a piece of evidence that fitted into an already existing hypothesis of the naturalness of homosexuality. Indeed, articles arguing for the naturalness and innateness of homosexuality took up the largest number of pages in Wilhelm’s review section, whereas the review section discussing homosexuality as acquired received far fewer contributions (Dubout, 2016: 60). The argument that homosexuality was inborn had long and deep roots: Karl Heinrich Ulrichs had already made the same claim several decades earlier (Ulrichs, 1870). Magnus Hirschfeld, too, was committed to finding scientific proof that homosexuality was inborn, and used this as a foundational argument for his gay rights activism. Hirschfeld’s double effort as initiator of sexological theories and methodologies, as well as gay rights activist, caused clashes with sexologists and scientists who proposed a different model of homosexuality, for example one based on sexual pathology.
What was particularly important about the argument that homosexuality was natural and inborn was that Wilhelm used this as proof that those who attacked Hirschfeld’s scientific endeavours as politically driven were in fact politically driven themselves. In another article, published in the
This move to depoliticise nature and instead cast a light on the retrograde politics of those who considered homosexuality to be pathological or acquired appeared in another review, in which Wilhelm criticised an article published by Gustav Fritsch, an anatomist, anthropologist, and neuropsychiatrist based in Berlin, on the ‘so-called third sex’ of humans. Fritsch, referring to both human and non-human animal organisms, argued that two sexes always appeared fully differentiated, and that even in cases where sex organs appeared hermaphroditic, this did not prove a bisexual disposition. Rather than assume an originary bisexuality, Fritsch suggested we should simply speak of ‘monstrosities’, the term denoting an unnatural abnormality of growth, a deviation from the norm of male and female sex. Wilhelm’s review of Fritsch’s article followed the typical pattern of his Bibliography, as outlined by Dubout: He heavily criticised Fritsch’s article, which argued against the bisexual predisposition of humans, and instead proposed the usual argument that homosexuality and other sexually intermediate conditions were inborn (Dubout, 2016). In order to prove Fritsch wrong, Wilhelm enlisted Ernst Haeckel’s work on gonochorism (sexual separation into two distinct sexes) versus hermaphroditism, which showed the prevalence of hermaphroditism in the animal kingdom and in humans, to argue that the primary disposition was clearly hermaphroditic; Eugen Steinach’s work on gonadal transplantation experiments to argue that transplanted sex glands could bring out a change in sexual behaviour only if the predisposition of that sex was already present; and finally, Goldschmidt’s butterfly experiments, arguing that these confirmed once and for all the hermaphroditic early stages of both butterflies and humans. Here, he made particular reference to Goldschmidt’s article on human intersex, arguing that Goldschmidt’s research showed that a quantitative system of potency decided which characteristics were latent and which manifest. For Wilhelm, this had particularly clear consequences for understanding nature’s ‘intent’: The apparent proof, which Fritsch deduces from the supposed tendency of nature towards sexual differentiation, is fairly weak, because we do not know which tendency nature pursues and, vice versa, we can deduce from the hermaphroditic living things which, without a doubt, exist, that nature has a tendency to bring forth a so-called Third Sex for some metaphysical purpose. (Praetorius, 1919: 177)
In Goldschmidt’s butterfly experiments Wilhelm recognised a supporting argument for his ongoing research into the innateness and naturalness of homosexuality. In finding Goldschmidt’s experimental proof and listing it next to the work of Haeckel and Steinach, Wilhelm built a case for the naturalness of homosexuality. In doing so, Wilhelm depoliticised nature and instead revealed its interpreters, such as Kraepelin and Fritsch, as driven by political goals. Goldschmidt, who was not associated with the group of sexologists centred on the SHC or the Institute of Sexology, and who nonetheless argued that intersexuality occurred in nature, also served as a suitable politically neutral person to underline the point that Hirschfeld and Wilhelm, who argued against the decriminalisation of homosexuality, were not politically driven, but that Kraepelin and Fritsch were. As a result, for Wilhelm’s – and, by extension, Hirschfeld’s – sexological project, Goldschmidt’s butterflies were politically potent.
Heredity and psychology: Arthur Kronfeld
Arthur Kronfeld (1886–1941) was a psychiatrist and founding member of the Institute of Sexology in Berlin, where he worked from 1919 to 1926 (Sigusch, 2008: 359). In 1922, the
In his article, Kronfeld listed what he considered to be two recent scientific developments that contributed to the understanding of the constitution of homosexuality: the discovery of hormones and the study of heredity. As Nelly Oudshoorn shows, once the concept of sex hormones had been developed after 1900, sex researchers quickly came forward to show the shared responsibility of hormones and genes for sexual development (Oudshoorn, 1994: 20–1). Sexual development itself was broken down into two stages: that of sex determination, demarcated as the realm of genetics, and that of sexual differentiation, the realm of endocrinology (ibid.: 21). Kronfeld’s interest in both hormones and the study of heredity therefore indicated a holistic interest in sexual development more broadly, rather than sex determination or differentiation specifically.
Kronfeld went on to argue that
Unlike Hodann and Wilhelm, Kronfeld displayed a much more critical stance on Hirschfeld’s theory of sexual intermediacy. Kronfeld added that most cases of homosexuality could not be matched exactly to laws of nature discovered by constitutional biology, because homosexuality stood in intimate relation to an individual’s psychological and social personality. Kronfeld was not entirely satisfied with Hirschfeld’s theory, because he considered it to be too biologistic (Herzer, 2017: 301). In his
In the conclusion to his article, Kronfeld argued that ‘same-sex-ness is connected with the very being of its carriers, it is fatefully connected with its constitution; it is not a coincidental perversion of the soul and the drives…but is a necessary and deep-seated desire of being [
As in the work of Hodann and Wilhelm, sexual intermediacy here was linked to the constitutional core of one’s being, but unlike Hodann and Wilhelm, Kronfeld proposed other equally important factors that influenced sexual development. This approach was only partially reconcilable with a sexual rights programme that had been established on the very basis of biological explanations for homosexuality. As a consequence, Kronfeld’s understanding of sexuality, which clearly acknowledged the constitutional force of an individual, but also sought to explore psychological factors, and did not give the same amount of credit to Goldschmidt’s butterflies as did Wilhelm and Hodann, would eventually lead to a break from Hirschfeld. In 1926, Kronfeld left the Institute of Sexology and opened his own psychiatric practice in Berlin.
The laws of nature and their limits: Arthur Weil
In 1923, Arthur Weil (1887–1969) published an article in the
Weil began his article by outlining the recent contributions of research in genetics to determine the inheritance of sex. Whilst experiments were often based on animal model organisms, Weil argued that the results of these experiments were relevant to the study of the human
Goldschmidt’s butterflies, Weil asserted, served as partial, but foundational evidence to uncover the laws of nature that determined the development of sex/sexuality in humans. Yet this reference to the natural laws, or laws of nature, is somewhat opaque, because it is not entirely clear whether Weil was using ‘nature’ to mean non-human animals, or individuals (whether human or non-human) ‘in the wild’ (as opposed to the laboratory). ‘In nature’, Weil argued, ‘one can often find, for example in butterflies such intersex forms’ (Weil, 1923: 29). Yet these intersex butterflies, which Weil claimed were to be found in nature, were actually found in Goldschmidt’s controlled laboratory environment. Weil referred to the ‘nature experiment’ – Goldschmidt’s butterflies – to argue that in the case of the butterfly, hormones played no important role. ‘Nature’ here referred to intersex butterflies bred in the carefully controlled space of the laboratory; what was natural about them was their animal biology. The laws of nature Weil sought to uncover, then, applied to non-human animals fully, but humans only partially, because of the ‘Höherentwicklung’ (‘higher development’ in evolutionary terms) of the latter (ibid.).
In a way similar to Kronfeld, Weil argued that butterfly experiments could illuminate the very core and origin of the human
Nature and artificiality: Magnus Hirschfeld
In 1923, Magnus Hirschfeld (1868–1935) published an article in the When I published the first volume of the
By pointing out that his own research had taken place before the major scientific works on the study of sex had been produced, Hirschfeld was situating his own sexological work at the avant-garde of sex research, and claiming that the very idea that sex might be worth studying was only just coming into being. Hirschfeld claimed that his own work recognised intersex before the topic was discussed, before research in biology, psychoanalysis, genetics, zoology, and endocrinology had come to the same conclusions, and before sexology had started to be taken seriously as a discipline. Hirschfeld thus subsumed the studies of a whole generation of scientists from diverse disciplines and their scientific evidence to the sexological project.
Hirschfeld’s speech-cum-article continued: When I drew up the study of sexual intermediate stages, I based this on purely clinical observations, but drawing on the broadest possible mass material, which did not come to me, but which I sought out. Each patient file collected in a consultation, even that of the most prolific psychiatrist, only shows a one-sided view. And as important as the findings of artificial breeding of bugs and butterflies may be, which have caused a deserved stir amongst geneticists, by far more value comes from studying the results of breeding which nature herself has done. (Hirschfeld, 1923: 9)
In contrast to the sociological study of sexual intermediate life, Hirschfeld implied, Goldschmidt’s butterflies lived, bred, and died as experimental objects in the lab, and the conditions of their life should therefore be considered less natural and of lower scientific value. Goldschmidt’s experimental studies, which were ‘artificial’, provided useful scholarship to draw upon, but these artificial experiments needed to be advanced through sexological observation ‘in nature’. Here, Hirschfeld’s approach showed a significant difference from Weil’s understanding of nature as animal biology. Hirschfeld's argument also exhibited an interesting inversion of the natural and the artificial. Hirschfeld saw a hierarchy between the study of heredity by way of experiments and that by way of clinical observation, a hierarchy or rivalry that, for example, Hodann did not recognise in his article from 1917.
Yet Hirschfeld clearly did find experiments on butterflies useful. In a report on the activities at the Institute during its first year in 1919, the following statement appears: During its first year, the actual experimental work did not advance, due to the difficult external conditions.…We also began the breeding of exotic silk-moths (

‘Schmetterlingszwitter’ (hermaphroditic butterflies; Hirschfeld, 1930: plate 33).
Conclusion: ‘The image of a butterfly coming to life’
In this article, I have explored how Goldschmidt’s experimental work on intersex butterflies was marshalled as evidence to illuminate the nature and naturalness of ‘sexual intermediacy’, in particular homosexuality, by sexological writers between 1917 and 1923, and how butterfly evidence informed the sexual politics of the group of sexologists affiliated with the Institute of Sexology. Hirschfeld’s sexology exchanged concepts with cutting-edge life sciences. In considering the history of the specific significance of intersex butterfly experiments in the sexological context, this article was not intended to judge retrospectively the successes or failures of certain scientific claims, but to show that this interdisciplinary interest that extended to neighbouring disciplines and their research on non-human animals was an integral part of sexology’s public profile during the interwar era. This history speaks to sexology’s dual life as a set of authoritative pronouncements (backed by biological expertise) and a countercultural endeavour to undercut existing norms.
Intersex butterflies functioned as important and politically charged evidence for a discussion at the heart of the sexological project of those involved in the founding of the Institute of Sexology: the question of the nature and naturalness of homosexuality (and sexual intermediacy more broadly) and its political consequences. Goldschmidt’s butterflies fluttered through the work of each of the sexologists discussed here, and provided evidence for their various theories. Their significance is far from obvious. Butterfly evidence showed a level of plasticity, as each sexologist incorporated Goldschmidt’s butterflies in different ways and in different stages of human sexual development: Hodann used the results of butterfly experiments to provide clinical applications; Wilhelm depoliticised nature and firmly ascribed bad politics to Hirschfeld’s opponents; Kronfeld only partially accepted their authority, but pushed beyond constitution to psychology; Weil looked to butterflies to understand universal laws of nature; and Hirschfeld criticised the artificiality of the butterfly experiment. At the same time, the use of butterfly evidence in the work of the sexologists discussed above shared a common reference point: the idea that non-human animals and humans were connected through a shared investment in a concept of sexual nature. Each of the intellectual intermediaries discussed in this article accepted that intersex butterflies could tell them something about human sexual variation. Goldschmidt’s intersex butterflies provided a potent tool for understanding sexuality in the human realm and for drawing sexual-political conclusions. These butterflies were also political actors in the history of sexology, offering a way of figuring out where people’s allegiances lay and how they fitted into a specific sexological vision that argued for a biologistic model of sexual intermediacy.
Although the importance of these butterflies for sexology has so far been neglected in scholarship on the history of sexology, traces of these butterflies remain. From 1919 to 1933, the Institute of Sexology stood at the corner of Beethovenstraße 3 and In den Zelten 10, close to the river Spree and in the heart of Berlin. After the Institute was ransacked in 1933 and destroyed in World War II, a conference centre was built on the site. Today, this is the Haus der Kulturen der Welt (HKW), the House of World Cultures. Here, in the vicinity of Hirschfeld’s former Institute, a sculpture by Henry Moore was erected in 1987 (Figure 4). The sculpture, a large bronze-coloured object positioned in the middle of a shallow pond, is called ‘Large Divided Oval: Butterfly’. The website of the HKW describes it thus:
The play of light and reflective water makes the sculpture appear as if it is opening upwards and expanding beyond the borders of its form. It plays with the impression of being a pair of wings, thus awakening most intriguingly the image of a butterfly coming to life.

‘Large Divided Oval: Butterfly’, Henry Moore, 1987, Photo: Sebastian Bolesch/HKW, 2010 (Haus der Kulturen der Welt, n. d.).
The presence of Moore’s butterfly sculpture where the Institute of Sexology once stood is a coincidence, but a fateful one. After the rise of National Socialism in Germany and the destruction of the Institute, German sexology ceased to exist in the way it had at the height of its success, and sexological research shifted to different geographic regions (as Katie Sutton’s article in this special issue shows). It was not until the 1980s that the history of sexology in Germany was taken up as a subject of study, most notably through the Magnus Hirschfeld Gesellschaft in West Berlin. Since then, the history of sexology has become a topic of increased interest and has, as this special issue shows, diversified into multiple histories of sexology. It is hoped that this article has contributed another history of sexology, one that ‘awaken[s] most intriguingly the image of a butterfly coming to life’ and makes a case for paying attention to non-human actors in the interdisciplinary history of sexology.
Footnotes
Declaration of conflicting interests
The author declared no potential conflicts of interest with respect to the research, authorship, and/or publication of this article.
Funding
The author disclosed receipt of the following financial support for the research, authorship, and/or publication of this article: Research for this article was funded by the Wellcome Trust under grant numbers 106653/Z/14/Z and 106654/Z/14/Z. In compliance with Open Data requirements, I declare that the research data supporting this publication are provided within this paper.
Notes
I presented the early stages of this work at the British Animal Studies Network meeting on the topic ‘sex’ at the University of Strathclyde and the Animals and Society Institute at the University of Illinois at Urbana-Champaign, and am grateful for the comments and feedback that I received there. I am extremely grateful to my anonymous reviewers and to the special issue editors, Katie Sutton and Kirsten Leng, for their careful reviews of this article. Their comments and feedback greatly improved my argument. I would like to thank the Rethinking Sexology team (Kate Fisher, Jana Funke, Jen Grove, Sarah Jones) and Kazuki Yamada for their comments on this article and support throughout. I also thank the Haus der Kulturen der Welt for permission to reproduce
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